Within undefined and no doubt variable geographical limits palaeontological zones have been found to be remarkably persistent. They follow each other in the same general order, but not always with equal definiteness. The type fossil may appear in some districts on a higher or a lower platform than it does in others. Only to a limited degree is there any coincidence between lithological variations in the strata and the sequence of the zones. In the Jurassic formations, indeed, where frequent alternations of different sedimentary materials are to be met with, it is in some cases possible to trace a definite upward or downward limit for a zone by some abrupt change in the sedimentation, such as from limestone to shale. But such a precise demarcation is impossible where no distinct bands of different sediments are to be seen. The zones can then only be vaguely determined by finding their characteristic fossils, and noting where these begin to appear in the strata and where they cease. It would seem, therefore, that the sequence of palaeontological zones, or life-horizons, has not depended merely upon changes in the nature of the conditions under which the organisms lived. We should naturally expect that these changes would have had a marked influence; that, for instance, a difference should be perceptible between the character of the fossils in a limestone and that of those in a shale or a sandstone. The environment, when a limestone was in course of deposition, would generally be one of clear water, favourable for a more vigorous and more varied fauna than where a shale series was accumulating, when the water would be discoloured, and only such animals would continue to live in it, or on the bottom, as could maintain themselves in the midst of mud. But no such lithological reason, betokening geographical changes that would affect living creatures, can be adduced as a universally applicable explanation of the occurrence and limitation of palaeontological zones. One of these zones may be only a few inches, or feet or yards in vertical extent, and no obvious lithological or other cause can be seen why its specially characteristic fossils should not be found just as frequently in the similar strata above and below. There is often little or no evidence of any serious change in the conditions of sedimentation, still less of any widespread physical disturbance, such as the catastrophes by which the older geologists explained the extinction of successive types of life.