It has been suggested that, where the life-zones are well defined, sedimentation has been extremely slow, and that though these zones follow each other with no break in the sedimentation, they were really separated by prolonged intervals of time during which organic evolution could come effectively into play. But it is not easy to explain how, for example in the Lower Lias, there could have been a succession of prodigious intervals, when practically no sediment was laid down, and yet that the strata should show no sign of contemporaneous disturbance or denudation, but succeed each other as if they had been accumulated by one continuous process of deposit. It must be admitted that the problem of life-zones in stratigraphical geology has not yet been solved. As Darwin first cogently showed, the history of life has been very imperfectly registered in the stratified parts of the earth's crust. Apart from the fact that, even under the most favourable conditions, only a small proportion of the total flora and fauna of any period would be preserved in the fossil state, enormous gaps occur where no record has survived at all. It is as if whole chapters and books were missing from a historical work. Some of these lacunae are sufficiently obvious. Thus, in some cases, powerful dislocations have thrown considerable portions of the rocks out of sight. Sometimes extensive metamorphism has so affected them that their original characters, including their organic contents, have been destroyed. Oftenest of all, denudation has come into play, and vast masses of fossiliferous rock have been entirely worn away, as is demonstrated by the abundant unconformabilities in the structure of the earth's crust.