In other Gregarines, however, those, namely, which pass inwards,
ultimately becoming "coelomic," as well as those which become entirely
intracellular, no epimerite is ever developed, and, further, the body
remains single or unseptate. These forms, which include, for instance,
_Monocystis_ (fig. 4), _Lankesteria_, _Diplocystis_, are
distinguished, as _Acephalina_ or _Aseptata_ (_Haplocyta_,
_Monocystida_), according to which character is referred to, from the
others, termed _Cephalina_ or _Septata_ (_Polycystida_).
The two sets of terms are not, however, completely identical or
interchangeable, for there are a few forms which possess an epimerite,
but which lack the division into protomerite and deutomerite, and are
hence known as _Pseudomonocystida_; this condition may be primitive
(_Doliocystis_) or (possibly) secondary, the partition having in
course of time disappeared. Again, _Stenophora_ is a septate form
which has become, secondarily, completely intracellular during the
young stages, and, doubtless correlated with this, shows no sign of an
epimerite.
[Illustration: From Wasielewski, after Leger.
FIG. 7.--Forms of Epimerites.
1, _Gregarina longa_.
2, _Sycia inopinata_.
3, _Pileocephalus heerii_.
4, _Stylorhynchus longicollis_.
5, _Beloides firmus_.
6, _Cometoides crinitus_.
7, _Geneiorhynchus monnieri_.
8, _Echinomera hispida_.
9, _Pterocephalus nobilis_.]
With regard to the epimerites themselves, they are of all variety of
form and shape and need not be described in detail (fig. 7). In one or
two cases, however, another variety of attaching organella is met
with. Thus in _Pterocephalus_, only the rostrum of the sporozoite
penetrates into the host-cell, and no epimerite is formed. Instead, a
number of fine root-like processes are developed from near the
anterior end, which pass in between the host-cells (fig. 5) and thus
anchor the parasite firmly. Similarly, in the curious
_Schizogregarinae_, the anterior end of the (unseptate) body forms a
number of stiff, irregular processes, which perform the same function
(fig. 8). It is to be noted that these processes are non-motile, and
not in any way comparable to pseudopodia, to which they were formerly
likened.
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