_Orders of Hexapoda._--In the present article it is only possible to
treat of the division of the Hexapoda into orders and sub-orders and of
the relations of these orders to each other. For further classificatory
details, reference must be made to the special articles on the various
orders. As regards the vast majority of insects, the orders proposed by
Linnaeus are acknowledged by modern zoologists. His classification was
founded mainly on the nature of the wings, and five of his orders--the
Hymenoptera (bees, ants, wasps, &c.), Coleoptera (beetles), Diptera
(two-winged flies), Lepidoptera (moths and butterflies), and Hemiptera
(bugs, cicads, &c.)--are recognized to-day with nearly the same limits
as he laid down. His order of wingless insects (Aptera) included
Crustacea, spiders, centipedes and other creatures that now form classes
of the Arthropoda distinct from the Hexapoda; it also included Hexapoda
of parasitic and evidently degraded structure, that are now regarded as
allied more or less closely to various winged insects. Consequently the
modern order Aptera comprises only a very small proportion of Linnaeus's
"Aptera"--the spring-tails and bristle-tails, wingless Hexapoda that
stand evidently at a lower grade of development than the bulk of the
class. The earwigs, cockroaches and locusts, which Linnaeus included
among the Coleoptera, were early grouped into a distinct order, the
Orthoptera. The great advance in modern zoology as regards the
classification of the Hexapoda lies in the treatment of a heterogeneous
assembly which formed Linnaeus's order Neuroptera. The characters of the
wings are doubtless important as indications of relationship, but the
nature of the jaws and the course of the life-history must be considered
of greater value. Linnaeus's Neuroptera exhibit great diversity in these
respects, and the insects included in it are now therefore distributed
into a number of distinct orders. The many different arrangements that
have been proposed can hardly be referred to in this article. Of special
importance in the history of systematic entomology was the scheme of F.
Brauer (1885), who separated the spring tails and bristle-tails as a
sub-class Apterygogenea from all the other Hexapoda, these forming the
sub-class Pterygogenea distributed into sixteen orders. Brauer in his
arrangement of these orders laid special stress on the nature of the
metamorphosis, and was the first to draw attention to the number of
Malpighian tubes as of importance in classification. Subsequent writers
have, for the most part, increased the number of recognized orders; and
during the last few years several schemes of classification have been
published, in the most revolutionary of which--that of A. Handlirsch
(1903-1904)--the Hexapoda are divided into four classes and thirty-four
orders! Such excessive multiplication of the larger taxonomic divisions
shows an imperfect sense of proportion, for if the term "class" be
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