allowed its usual zoological value, no student can fail to recognize
that the Hexapoda form a single well-defined class, from which few
entomologists would wish to exclude even the Apterygogenea. In several
recent attempts to group the orders into sub-classes, stress has been
laid upon a few characters in the imago. C. Borner (1904), for example,
considers the presence or absence of cerci of great importance, while F.
Klapalek (1904) lays stress on a supposed distinction between
appendicular and non-appendicular genital processes. A natural system
must take into account the nature of the larva and of the metamorphosis
in conjunction with the general characters of the imago. Hence the
grouping of the orders of winged Hexapoda into the divisions
Exopterygota and Endopterygota, as suggested by D. Sharp, is unlikely to
be superseded by the result of any researches into minute imaginal
structure. Sharp's proposed association of the parasitic wingless
insects in a group Anapterygota cannot, however, be defended as natural;
and recent researches into the structure of these forms enables us to
associate them confidently with related winged orders. The
classification here adopted is based on Sharp's scheme, with the
addition of suggestions from some of the most recent authors--especially
Borner and Enderlein.
Class: HEXAPODA.
Sub-class: APTERYGOTA.
Primitively (?) wingless Hexapods with cumacean mandibles, distinct
maxillulae, and locomotor abdominal appendages. Without ectodermal
genital ducts. Young closely resemble adults.
The sub-class contains a single
Order: _Aptera_,
which is divided into two sub-orders:
1. _Thysanura_ (Bristle-tails): with ten abdominal segments; number of
abdominal appendages variable. Cerci prominent. Developed tracheal
system.
2. _Collembola_ (Spring-tails): with six abdominal segments;
appendages of the first forming an adherent ventral tube, those of the
third a minute "catch," those of the fourth (fused basally) a
"spring." Tracheal system reduced or absent.
Sub-class: EXOPTERYGOTA.
Hexapoda mostly with wings, the wingless forms clearly degraded.
Maxillulae rarely distinct. No locomotor abdominal appendages. The
wing-rudiments develop visibly outside the cuticle. Young like or
unlike parents.
Order: _Dermaptera_.
Biting mandibles; minute but distinct-maxillulae; second maxillae
incompletely fused. When wings are present, the fore-wings are small
firm elytra, beneath which the delicate hind-wings are complexly
folded. Many forms wingless. Genital ducts entirely mesodermal. Cerci
always present; usually modified into unjointed forceps. Numerous (30
or more) Malpighian tubes. Young resembling parents.
Includes two families--the _Forficulidae_ or _earwigs_ (q.v.) and the
_Hemimeridae_.
Order: _Orthoptera_.
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