[Illustration: _Fig. 54._--Diagram showing possible modifications of
the persons of a Calyptoblastic Hydromedusa. Letters a to h same as in
fig. 49. i, The horny cup or hydrotheca of the hydriform persons; l,
medusiform person springing from m, a modified, hydriform person
(blastostyle); n, the horny case or gonangium enclosing the
blastostyle and its buds. This and the hydrotheca i give origin to the
name _Calyptoblastea_. (After Allman.)]
_Perigonimus._--This common British hydroid belongs by its characters
to the family _Bougainvillidae_; it produces, however, a medusa of the
genus _Tiara_ (fig. 52), referable to the family _Clavidae_; a fact
sufficient to indicate the tentative character of even the most modern
classifications of this order.
SUB-ORDER II. HYDROIDEA CALYPTOBLASTEA (LEPTOMEDUSAE).--Trophosome with
polyps always differentiated into nutritive and reproductive individuals
(blastostyles) enclosed in hydrothecae and gonothecae respectively; with
sympodial type of budding. Gonosome with free medusae or gonophores; the
medusae typically with otocysts, sometimes with cordyli or ocelli (figs.
54, 55).
[Illustration: FIG. 55.--View of the Oral Surface of one of the
_Leptomedusae_ (_Irene pellucida_, Haeckel), to show the numerous
tentacles and the otocysts.
ge, Genital glands.
M, Manubrium.
ot, Otocysts.
rc, The four radiating canals.
Ve, The velum.]
The calyptoblastic polyp of the nutritive type is very uniform in
character, its tendency to variation being limited, as it were, by the
enclosing hydrotheca. The hydranth almost always has a single circlet of
tentacles, like the _Bougainvillea_-type, in the preceding sub-order; an
exception is the curious genus _Clathrozoon_, in which the hydranth has
a single tentacle. The characteristic hydrotheca is formed by the bud at
an early stage (fig. 56); when complete it is an open cup, in which the
hydranth develops and can be protruded from the opening for the capture
of food, or is withdrawn into it for protection. Solitary polyps are
unknown in this sub-order; the colony may be creeping or arborescent in
form; if the latter, the budding of the polyps, as already stated, is of
the sympodial type, and either biserial, forming stems capable of
further branching, or uniserial, forming pinnules not capable of further
branching. In the biserial type the polyps on the two sides of the stem
have primitively an alternating, zigzag arrangement; but, by a process
of differential growth, quickened in the 1st, 3rd, 5th, &c., members of
the stem, and retarded in the 2nd, 4th, 6th, &c., members, the polyps
may assume secondarily positions opposite to one another on the two
sides of the stem. Other variations in the mode of growth or budding
bring about further differences in the building up of the colony, which
are not in all cases properly understood and cannot be described in
detail here. The stem may contain a single coenosarcal tube
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