("monosiphonic") or several united in a common perisarc
("polysiphonic"). An important variation is seen, in the form of the
hydrotheca itself, which may come off from the main stem by a stalk, as
in _Obelia_, or may be sessile, without a stalk, as in _Sertularia_.
[Illustration: After Allman, _Gymnoblastic Hydroids_, by permission of
the council of the Ray Society.
FIG. 56.--Diagrams to show the mode of formation of the Hydrotheca and
Gonotheca in Calyptoblastic Hydroids. A-D are stages common to both;
from D arises the hydrotheca (E) or the gonotheca (F); th, theca; st,
stomach; t, tentacles; m, mouth; mb, medusa-buds.]
In many Calyptoblastea there occur also reduced defensive polyps or
dactylozoids, which in this sub-order have received the special name of
_sarcostyles_. Such are the "snake-like zoids" of _Ophiodes_ and other
genera, and as such are generally interpreted the "machopolyps" of the
_Plumularidea_. These organs are supported by cuplike structures of the
perisarc, termed nematophores, regarded as modified hydrothecae
supporting the specialized polyp-individuals. They are specially
characteristic of the family _Plumularidae_.
The medusa-buds, as already stated, are always produced from
blastostyles, reduced non-nutritive polyps without mouth or tentacles.
An apparent, but not real, exception is _Halecium halecinum_, in which
the blastostyle is produced from the side of a nutritive polyp, and both
are enclosed in a common theca without a partition between them (Allman
[1] p. 50, fig. 24). The gonotheca is formed in its early stage in the
same way as the hydrotheca, but the remains of the hydranth persists as
an operculum closing the capsule, to be withdrawn when the medusae or
genital products are set free (fig. 56).
The blastostyles, gonophores and gonothecae furnish a series of
variations which can best be considered as so many stages of
evolution.
Stage 1, seen in _Obelia_. Numerous medusae are budded successively
within the gonotheca and set free; they swim off and mature in the
open sea (Allman [1], p. 48, figs. 18, 19).
Stage 2, seen in _Gonothyraea_. Medusae, so-called "meconidia," are
budded but not liberated; each in turn, when it reaches sexual
maturity, is protruded from the gonotheca by elongation of the stalk,
and sets free the embryos, after which it withers and is replaced by
another (Allman [1], p. 57, fig. 28).
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