2. _Stylasteridae._--Coenosteum arborescent, sometimes fanlike, with
pores only on one face, or on the lateral margins of the branches;
gastropores with tabulae only in two genera, but with (except in
_Astylus_) a _style_, i.e. a conical, thorn-like projection from the
base of the pore, sometimes found also in dactylopores; sessile
gonophores. _Sporadopora_ has the pores scattered irregularly.
_Distichopora_ has the pores arranged in rows. _Stylaster_ has
cyclosystems. In _Allopora_ the cyclostems resemble the calyces of
Anthozoan corals. In _Cryptohelia_ the cyclosystem is covered by a cap
or operculum. In _Astylus_ (fig. 63) styles are absent.
_Affinities of the Hydrocorallinae._--There can be no doubt that the
forms comprised in this order bear a close relationship to the
Hydroidea, especially the sub-order Gymnoblastea, with which they
should perhaps be classed in a natural classification. A
hydrocoralline may be regarded as a form of hydroid colony in which
the coenosarc forms a felt-work ramifying in all planes, and in which
the chitinous perisarc is replaced by a massive calcareous skeleton.
So far as the trophosome is concerned, the step from an encrusting
hydroid such as _Hydractinia_ to the hydrocoralline _Millepora_ is not
great.
Hickson considers that the families _Milleporidae_ and _Stylasteridae_
should stand quite apart from one another and should not be united in
one order. The nearest approach to the _Stylasteridae_ is perhaps to
be found in _Ceratella_, with its arborescent trophosome formed of
anastomosing coenosarcal tubes supported by a thick perisarc and
covered by a common ectoderm. _Ceratella_ stands in much the same
relation to the _Stylasteridae_ that _Hydractinia_ does to the
_Milleporidae_, in both cases the chitinous perisarc being replaced by
the solid coenosteum to which the hydrocorallines owe the second half
of their name.
ORDER IV. Graptolitoidea (Rhabdophora, Allman).--This order has been
constituted for a peculiar group of palaeozoic fossils, which have been
interpreted as the remains of the skeletons of Hydrozoa of an extinct
type.
A typical graptolite consists of an axis bearing a series of tooth-like
projections, like a saw. Each such projection is regarded as
representing a cup or hydrotheca, similar to those borne by a
calyptoblastic hydroid, such as _Sertularia_. The supposed hydrothecae
may be present on one side of the axis only (monoprionid) or on both
sides (diprionid); the first case may be conjectured to be the result of
uniserial (helicoid) budding, the second to be produced by biserial
(scorpioid) budding. In one division (_Retiolitidae_) the axis is
reticulate. In addition to the stems bearing cups, there are found
vesicles associated with them, which have been interpreted as gonothecae
or as floats, that is to say, air-bladders, acting as hydrostatic organs
for a floating polyp-colony.
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