_Vespoidea._--This section includes a number of families characterized
by the backward extension of the prothorax to the tegulae and
distinguished from the ants by the absence of "nodes" at the base of
the abdomen. The true wasps have the fore-wings folded lengthwise when
at rest and the fore-legs of normal build--not specialized for
digging. The _Vespidae_ or social wasps have "queens" and "workers"
like the ants, but both these forms of female are winged; the claws on
their fret are simple. In the _Eumenidae_ or solitary wasps the female
sex is undifferentiated, and the foot claws are toothed. (For the
habits of these insects see WASP.) The _Chrysididae_ or ruby wasps are
small insects with a very hard cuticle exhibiting brilliant metallic
colours--blue, green and crimson. Only three or four abdominal
segments are visible, the hinder segments being slender and retracted
to form a telescope-like tube in which the ovipositor lies. When the
ovipositor is brought into use this tube is thrust out. The eggs are
laid in the nests of various bees and wasps, the chrysid larva living
as a "cuckoo" parasite. The _Trigonalidae_, a small family whose
larvae are parasitic in wasps' nests, also probably belong here.
The other families of the _Vespoidea_ belong to the series of
"Fossores" or digging-wasps. In two of the families--the _Mutillidae_
and _Thynnidae_--the females are wingless and the larvae live as
parasites in the larvae of other insects; the female _Mutilla_ enters
bumble-bees' nests and lays her eggs in the bee-grubs. In the other
families both sexes are winged, and the instinct and industry of the
females are among the most wonderful in the Hymenoptera. They make
burrows wherein they place insects or spiders which they have caught
and stung, laying their eggs beside the victim so that the young
larvae find themselves in presence of an abundant and appropriate
food-supply. Valuable observations on the habits of these insects are
due to J. H. Fabre and G. W. and E. Peckham. The prey is sometimes
stung in the neighbourhood of the nerve ganglia, so that it is
paralysed but not killed, the grub of the fossorial wasp devouring its
victim alive; but this instinct varies in perfection, and in many
cases the larva flourishes equally whether its prey be killed or not.
The females have a wonderful power of finding their burrows on
returning from their hunting expeditions. Among the Vespoid families
of fossorial wasps, the _Pompilidae_ are the most important. They are
recognizable by their slender and elongate hind-legs; many of them
provision their burrows with spiders. The _Sapygidae_ are parasitic on
bees, while the _Scoliidae_ are large, robust and hairy insects, many
of which prey upon the grubs of chafers.
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