_Sphecoidea._--In this division are included the rest of the
"digging-wasps," distinguished from the _Vespoidea_ by the short
pronotum not reaching backward to the tegulae. They have usually been
reckoned as forming a single, very large family--the _Sphegidae_--but
ten or twelve subdivisions of the group are regarded as distinct
families by Ashmead and others. Great diversity is shown in the
details of structure, habits and nature of the prey. Species of
_Sphex_, studied by Fabre, provisioned their brood-chambers with
crickets. _Pelopoeus_ hunts spiders, while _Ammophila_ catches
caterpillars for the benefit of her young. Fabre states that the
last-named insect uses a stone for the temporary closing of her
burrow, and the Peckhams have seen a female _Ammophila_ take a stone
between her mandibles and use it as a hammer for pounding down the
earth over her finished nest. The habits of _Bembex_ are of especial
interest. The female, instead of provisioning her burrow with a supply
of food that will suffice the larva for its whole life, brings fresh
flies with which she regularly feeds her young. In this instinct we
have a correspondence with the habits of social wasps and bees. Yet it
may be thought that the usual instinct of the "digging-wasps" to
capture and store up food in an underground burrow for the benefit of
offspring which they will never see is even more surprising. The habit
of some genera is to catch the prey before making their tunnel, but
more frequently the insect digs her nest, and then hunts for prey to
put into it.
_Apoidea._--The bees which make up this group agree with the
Sphecoidea in the short pronotum, but may be distinguished from all
other Hymenoptera by the widened first tarsal segment and the plumose
hairs on head and body. They are usually regarded as forming a single
family--the _Apidae_--but there is very great diversity in structural
details, and Ashmead divides them into fourteen families. The
"tongue," for example, is short and obtuse or emarginate in _Colletes_
and _Prosopis_, while in all other bees it is pointed at the tip. But
in _Andrena_ and its allies it is comparatively short, while in the
higher genera, such as _Apis_ and _Bombus_, it is elongate and
flexible, forming a most elaborate and perfect organ for taking liquid
food. Bees feed on honey and pollen. Most of the genera are "solitary"
in habit, the female sex being undifferentiated; but among the
humble-bees and hive-bees we find, as in social wasps and ants, the
occurrence of workers, and the consequent elaboration of a wonderful
insect-society. (See BEE.)
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