It has been seen that the Flemish school of illumination in the 13th and 14th centuries followed the French model. In the 15th century, while the old tradition continued in force for a while, the art developed on an independent line; and in the second half of the century it exercised a widespread influence on the neighbouring countries, on France, on Holland and on Germany. This development was one of the results of the industrial and artistic activity of the Low Countries at this period, when the school of the Van Eycks and their followers, and of other artists of the great and wealthy cities, such as Bruges, Antwerp, Ghent, were so prolific. The Flemish miniatures naturally followed on the lines of painting. The new style was essentially modern, freeing itself from the traditions of medieval illumination and copying nature. Under the hand of the Flemish artist the landscape attained to great perfection, softness and depth of colouring, the leading attribute of the school, lending a particular charm and sense of reality to his out-door scenes. His closer observation of nature is testified also in the purely decorative part of his work. Flowers, insects, birds and other natural objects now frequent the border, the origin of which is finally forgotten. It ceases to be a connected growth wandering round the page; it becomes a flat frame of dull gold or colour, over which isolated objects, flowers, fruits, insects, butterflies, are strewn, painted with naturalistic accuracy and often made, by means of strong shadows, to stand out in relief against the background. This practice was soon carried to florid excess, and all kinds of objects, including jewels and personal ornaments, were pressed into the service of the border, in addition to the details copied from nature. The soft beauty of the later Flemish style proved very attractive to the taste of the day, with the result that it maintained a high standard well on into the 16th century, the only rivals being the MSS. of Italian art. The names of celebrated miniaturists, such as Memlinc, Simon Bening of Ghent, Gerard of Bruges, are associated with its productions; and many famous extant examples bear witness to the excellence to which it attained. The Grimani Breviary at Venice is one of the best known MSS. of the school; but almost every national library has specimens to boast of. Among those in the British Museum may be mentioned the breviary of Queen Isabella of Spain (Add. MS. 18,851); the Book of Hours of Juana of Castille (Add. MS. 18,852); a very beautiful Book of Hours executed at Bruges (Egerton MS. 2125); another exquisite but fragmentary MS. of the same type (Add. MS. 24,098) and cuttings from a calendar of the finest execution (Add. MS. 18,855) ascribed to Bening of Ghent; a series of large sheets of genealogies of the royal houses of Portugal and Spain (Add. MS. 12,531) by the same master and others; and late additions to the Sforza Book of Hours (Add. MS. 34,294).