A characteristic feature of glandular activity is the production of a
slight electrical disturbance. In the case of _Malopterurus_ this
elsewhere subsidiary function of the skin has become so exaggerated as
to lead to the conversion of the skin of each side of the body into a
powerful electrical organ.[3] Each of these consists of some two
million small chambers, each containing an electric disk and all
deriving their nerve supply from the branches of a single enormous
axis cylinder. This takes its origin from a gigantic ganglion cell
situated latero-dorsally in the spinal cord between the levels of the
first and second spinal nerves.
_Cement Organs._--The larvae of certain Teleostomes and Dipnoans possess
special glandular organs in the head region for the secretion of a
sticky cement by which the young fish is able to attach itself to
water-plants or other objects. As a rule these are ectodermal in origin;
e.g. in _Lepidosiren_ and _Protopterus_[4] the crescentic cement organ
lying ventrally behind the mouth consists of a glandular thickening of
the deep layer of the ectoderm. In young ganoid fishes preoral cement
organs occur. In Crossopterygians there is one cup-shaped structure on
each side immediately in front of the mouth. Here the glandular
epithelium is endodermal, developed[5] as an outgrowth from the wall of
the alimentary canal, closely resembling a gill pouch. In _Amia_[6] the
same appears to be the case. In a few Teleosts similar organs occur,
e.g. _Sarcodaces_, _Hyperopisus_,[7] where so far as is known they are
ectodermal.
_Photogenic Organs._--The slimy secretion produced by the epidermal
glands of fishes contains in some cases substances which apparently
readily undergo a slow process of oxidation, giving out light of low
wave-length in the process and so giving rise to a phosphorescent
appearance. In many deep-sea fishes this property of producing
light-emitting secretion has undergone great development, leading to the
existence of definite photogenic organs. These vary much in character,
and much remains to be done in working out their minute structure. Good
examples are seen in the Teleostean family _Scopelidae_, where they form
brightly shining eye-like spots scattered about the surface of the body,
especially towards the ventral side.
[Illustration: From _Trans. Zool. Soc. of London_.
FIG. 6.--Larva of Polypterus. (After Budgett.)]
[Illustration: From _Phil. Transactions, Royal Society of London_.
FIG. 7.--Thirty Days' Larval Lepidosiren. (After Graham Kerr.)]
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