to them, has been expounded by Professor Giard, namely, that it tends
to obliterate the secondary sexual characters. Modern research has
discovered no crab to surpass _Macrocheira kämpferi_, De Haan, that
can span between three and four yards with the tips of its toes, but
at the other end of the scale it has yielded _Collodes malabaricus_,
Alcock, "of which the carapace, in an adult and egg-laden female, is
less than one-sixth of an inch in its greatest diameter." The most
abyssal of all crabs yet known is _Ethusina abyssicola_, Smith, or
what is perhaps only a variety of it, _E. challengeri_, Miers. Of the
latter the "Albatross" obtained a specimen from a depth of 2232
fathoms (Faxon, 1895), of the former from 2221 fathoms, and of this S.
I. Smith remarks that it has "distinctly faceted black eyes," although
in them "there are only a very few visual elements at the tips of the
immobile eye-stalks."
The _Brachyura anomala_, or Dromiidea, "have preserved the external
characters and probably also the organization of the Brachyura of the
Secondary epoch" (Milne-Edwards and Bouvier, 1901). They agree with
the true crabs in not having appendages (uropods) to the sixth segment
of the pleon, the atrophy being complete in the Homolidae and
Homolodromiidae, whereas in the Dromiidae and Dynomenidae a pair of
small plates appear to be vestiges of these organs. In the family
Homolidae stands the strange genus _Latreillia_, Roux, with long
slender limbs and triangular carapace after the fashion of oxyrhynch
spider-crabs. In _Homola_ the carapace is quadrilateral. Between these
two a very interesting link was discovered by the "Challenger" in the
species _Latreillopsis bispinosa_. Henderson. Bouvier (1896) has shown
that _Palaeinachus longipes_, Woodward, from the Forest Marble of
Wiltshire, is in close relationship, not to the oxyrhynch Inachidae,
but to the genera _Homolodromia_ and _Dicranodromia_ of the
Homolodromiidae, and that the Jurassic crabs in general, of the family
Prosoponidae (Meyer), are Dromiidea.
Public-domain text, read in full here on John Shaqi.
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