2. MACRURA.--The _Macrura anomala_, or Anomura in restricted sense,
are popularly known through the hermit-crabs alone. These only
partially represent one of the three main divisions, Paguridea,
Galatheidea, Hippidea. The first of these is subdivided into
_Pagurinea_, _Lithodinea_, _Lomisinea_, each with a literature of its
own. Among the Pagurinea is the _Birgus latro_, or robber-crab, whose
expertness in climbing the coco-nut palm need no longer be doubted,
since in recent years it has been noted and photographed by
trustworthy naturalists in the very act. Alcock "observed one of these
crabs drinking from a runnel of rain-water, by dipping the fingers of
one of its chelipeds into the water and then carrying the wet fingers
to its mouth." Hermits of the genus _Coenobita_ he found feeding
voraciously on nestling sea-terns. That pagurids must have the usually
soft pleon or abdomen protected by the shell of a mollusc is now known
to be subject to a multitude of exceptions. _Birgus_ dispenses with a
covering; _Coenobita_ can make shift with half the shell of a
coco-nut; _Chlaenopagurus_ wraps itself up in a blanket of colonial
polyps; _Cancellus tanneri_, Faxon, was found in a piece of dead coral
rock; _Xylopagurus rectus_, A. Milne-Edwards, lodges in tubes of
timber or bits of hollow reed. The last-named species has a straight
symmetrical abdomen, with the penultimate segment expanded and
strongly calcified to form a back-door to the very unconventional
habitation. This it enters head-foremost from the rear, while
"hermits" in general are forced to go backwards into their spiral or
tapering shelters by the front. Some of the species can live in the
ocean at a depth of two or three miles. Some can range inland up to a
considerable height on mountains. The advantage that this group has
derived from the adoption of mollusc shells as houses or fortresses,
ready built and light enough for easy transport, is obviously
discounted by a twofold inconvenience. There is nothing to ensure that
the supply will be equal to the demand, and Nature has not arranged
that the borrowed tenement shall continue to grow with the growth of
its new tenant. To meet these defects it is found that numerous
species encourage or demand the companionship of various zoophytes,
simple or colonial. These sometimes completely absorb the shell on
which they are settled, but then act as a substitute for it, and in
any case by their outgrowth they extend the limits of the dwelling, so
that the inmate can grow in comfort without having to hunt or fight
for a larger abode. Among the _Lithodinea_, or stone crabs, besides
important readjustments of classification (Bouvier, 1895, 1896),
should be noticed the evidence of their cosmopolitan range, and the
species _Neolithodes agassizii_ (Smith) and _N. grimaldii_,
Milne-Edwards and Bouvier, which carry to an extreme the spinosity
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