5. SYMPODA.--This order of sessile-eyed decapods was absolutely
unknown to science till 1779. A species certainly belonging to it was
described by Lepekhin in 1780, but the obscure _Gammarus esca_, "food
Gammarus" beloved of herrings, described by J. C. Fabricius in the
preceding year, may also be one of its members. Nutritious
possibilities are implied in _Diastylis rathkii_, Kröyer, one of the
largest forms, which, though slender and rarely an inch long, in its
favourite Arctic waters is found "in incalculable masses, in thousands
of specimens" (Stuxberg, 1880). Far on in the 19th century eminent
naturalists were still debating whether in this group there were eyes
or no eyes, whether the eyes were stalked or sessile, whether the
animals observed were larval or adult. The American T. Say in 1818
gave a good description of a new species and founded the premier genus
_Diastylis_, but other investigators derived little credit from the
subject till more than sixty years after its introduction by the
Russian Lepekhin. Then Goodsir, Kröyer, Lilljeborg, Spence Bate and
one or two others made considerable advances, and in 1865 a memorable
paper by G. O. Sars led the way to the great series of researches
which he has continued to the present day. The name _Cumacea_,
however, which he uses cannot be retained, being founded on the
preoccupied name _Cuma_ (Milne-Edwards, 1828). The more recent name
_Sympoda_ (see Willey, _Results_, pt. v. p. 609, 1900) alludes to the
huddling together of the legs, which is conspicuous in most of the
species. Ten families are now distinguished--Diastylidae, Lampropidae,
Platyaspidae, Pseudocumidae, all with an articulated telson; without
one, the Bodotriidae (formerly called _Cumidae_), Vaunthompsoniidae,
Leuconidae, Nannastacidae, Campylaspidae, Procampylaspidae. All the
Leuconidae and Procampylaspidae are blind, and some species in most of
the other families. Usually the sides of the carapace are strangely
produced into a mock rostrum in front of the ocular lobe, be it
oculiferous or not. The last four or five segments of the trunk are
free from the carapace. The slender pleon has always six distinct
segments, the sixth carrying two-branched uropods, the preceding five
armed with no pleopods in the female, whereas in the male the number
of pairs varies from five to none. The resemblance of these creatures
to miniature Macrura is alluded to in the generic name _Nannastacus_,
meaning dwarf-lobster. In this genus alone of the known Sympoda the
eyes sometimes form a pair, in accordance with the custom of all other
malacostracan orders except this and of this order itself in the
embryo (Sars, 1900). The most but not the only remarkable character
lies in the first maxillipeds. These, with the main stem more or less
pediform, have the epipod and exopod modified for respiratory
purposes. The backward-directed epipods usually carry branchial
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account