vesicles. The forward-directed exopods either act as valves or form a
tube (rarely two tubes), protensile and retractile, for regulating
egress of water from the branchial regions. This mechanism as a whole
is unique, although, as Sars observes, the epipod of the first
maxillipeds has a respiratory function also in the Lophogastridae and
Mysidae and in the cheliferous isopods. As a rule armature of the
carapace is much more developed in the comparatively sedentary female
than in the usually more active male. Only in the male do the second
antennae attain considerable length, with strong resemblance to what
is found in some of the Amphipoda. About 150 species distributed among
thirty-four genera are now known, many from shallow water and from
between tide-marks, some from very great depths. H. J. Hansen
concludes that "they are all typically ground animals, and as yet no
species has been taken under such conditions that it could be reckoned
to the pelagic plankton." As they have been found in all zones and
chiefly by a very few observers, it is probable that a great many more
species remain to be discovered. In recent years thirteen species, all
belonging to the same genus _Pseudocuma_ (fig. 3), have been recorded
by Sars from the Caspian Sea. A bibliography of the order is given in
that author's _Crustacea of Norway_, vol. iii. (1899-1900).
[Illustration: Fig. 4.--_Rhabdosoma piratum_, Stebbing.]
6. ISOPODA.--This vast and populous order can be traced far back in
geological time. It is now represented in all seas and lands, in
fresh-water lakes and streams, and even in warm springs. It adapts
itself to parasitic life not only in fishes, but in its own class
Crustacea, and that in species of every order, its own included. In
this process changes of structure are apt to occur, and sometimes
unimaginable sacrifices of the normal appearance. The order has been
divided into seven tribes, of which a fuller summary than can here be
given will be found in Stebbing, _History of Crustacea_ (1893). The
first tribe, called Chelifera, from the usually chelate or
claw-bearing first limbs, may be regarded as _Isopoda anomala_, of
which some authors would form a separate order, Tanaidea. Like the
genuine isopods, they have seven pairs of trunk-legs, but instead of
having seven segments of the middle body (or peraeon) normally free,
they have the first one or two of its segments coalesced with the
head. Instead of the breathing organs being furnished by the
appendages of the pleon with the heart in their vicinity, the
respiration is controlled by the maxillipeds, with the heart in the
peraeon (see Delage, _Arch. Zool. expér. et gén._, vol. ix., 1881).
There are two families, Tanaidae and Apseudidae. Occasionally the
ocular lobes are articulated.
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