7. AMPHIPODA.--As in the genuine Isopoda, the eyes of Amphipoda are
always sessile, and generally paired, and, in contrast to crabs and
lobsters, these two groups have only four pairs of mouth-organs
instead of six, but seven pairs of trunk-legs instead of five. From
the above-named isopods the present order is strongly differentiated
by having heart and breathing organs not in the pleon, but in the
peraeon, or middle body, the more or less simple branchial vesicles
being attached to some or all of the last six pairs of trunk-legs.
Normally the pleon carries six pairs of two-branched appendages, of
which the first three are much articulated flexible swimming feet, the
last three few-jointed comparatively indurated uropods. There are
three tribes, _Gammaridea_, _Caprellidea_, _Hyperiidea_. The middle
one contains but two families, the cylindrical and often thread-like
skeleton shrimps, Caprellidae, and their near cousins, the broad,
flattened, so-called whale-lice, Cyamidae. This tribe has the pleon
dwindled into insignificance, whereas in the other two tribes it is
powerfully developed. The Hyperiidea are distinguished by having their
maxillipeds never more than three-jointed. In the companion tribes
these appendages have normally seven joints, and always more than
three. The order thus sharply divided is united by an intimate
interlacing of characters, and forms a compact whole at present
defying intrusion from any other crustacean group. Since 1775, when J.
C. Fabricius instituted the genus _Gammarus_ for five species, of
which only three were amphipods, while he left five other amphipods in
the genus _Oniscus_, from this total of eight science has developed
the order, at first very slowly, but of late by great leaps and
bounds, so that now the _Gammaridea_ alone comprise more than 1300
species, distributed among some 300 genera and 39 families. They
burrow in the sands of every shore; they throng the weeds between
tide-marks; they ascend all streams; they are found in deep wells, in
caverns, in lakes; in Arctic waters they swarm in numbers beyond
computation; they find lodgings on crabs, on turtles, on weed-grown
buoys; they descend into depths of the ocean down to hundreds or
thousands of fathoms; they are found in mountain streams as far above
sea-level as some of their congeners live below it. The Talitridae,
better known as sandhoppers, can forgo the briny shore and content
themselves with the damp foliage of inland forests or casual humidity
in the crater of an extinct volcano. Over the ocean surface, as well
as at various depths, float and swim innumerable _Hyperiidea_--the
wonderful _Phronima_, glass-like in its glassy barrel hollowed out of
some Tunicate; the _Cystisoma_, 4 or 5 in. long, with its eye-covered
head; the _Rhabdosoma_, like a thin rod of glass, with needle-like
head and tail, large eyes, but limbs and mouth-organs all in
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