Questions of fact are simple to deal with. It is indubitable that some
forms of life remain stationary and unevolving for secular periods; it
is equally indubitable that degeneration is widespread in evolution.
These are facts. But we are not therefore called upon to deny the
possibility of progress. To do so would be to fall into the error
of reasoning which we have already condemned. It remains for us to
take these facts into account when examining the totality of facts
concerning organic life, and to see whether, in spite of them, we
cannot discover a series of other facts, a movement in phenomena, which
may still legitimately be called progress. To deny progress because of
degeneration is really no more legitimate than to assert that, because
each wave runs back after it has broken, therefore the tide can never
rise.
Similarly with the first two objections. If the degree of adaptation
has not increased during evolution, then it is clear that progress
does not consist in increase in adaptation. But it does not follow
that progress does not exist; it may quite well consist in an increase
of other qualities. So with complexity. Complexity has increased, but
increase in complexity is not progress, say the objectors. Granted:
but may there not be something else which has increased besides mere
complexity?
No; the remedy for all our difficulties, and indeed the only way in
which we can arrive at the _possibility_ of saying whether biological
progress exists or no, is to adopt the positive method.
Let us then begin our survey of biological evolution in the endeavour
to find whether or no progress is visible there. To start with, we must
be clear what are the sources of our knowledge on the subject.
Direct observation of progressive evolution has, of course, not yet
been possible in the period--biologically negligible--in which man has
directed his attention to the problem; and historical record is also
absent. The best available evidence is that of paleontology: here the
relative positions of the layers of the earth’s crust enable us to
deduce their temporal sequence--and naturally, that of the organisms
whose fossil remains they embalm--with a great deal of accuracy.[3]
We can scarcely ever observe the direct transition from the forms of
life in an older to those in a younger stratum, nor can we absolutely
prove their genetic relationship. But in a vast number of cases it is
abundantly clear that the later type of organization is descended from
the former--that a group of forms in the younger stratum had its origin
in one or more species of the group to which the forms in the older
stratum belong. Sometimes, however, as in many groups of mammals, the
gaps are few and small, the seriation almost complete. In any event we
have here evidence which, so far as it goes, is perfectly admissible
for the main lines and for many of the smaller branches of evolutionary
descent.
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