Essays Upon Heredity and Kindred Biological Problems: Authorised TranslationWeismann, August
Science
Essays Upon Heredity and Kindred Biological Problems: Authorised Translation
Weismann, August
Evolution (Biology); Genetics; Reproduction
Sexual reproduction can also increase the differences between
individuals, because constant cross-breeding must necessarily and
repeatedly lead to a combination of forces which tend in the same
direction, and which may determine the constitution of any part of the
body. If, for instance, the same part of the body is strongly developed
in both parents, the experience of breeders tells us that the part in
question is likely to be even more strongly developed in the offspring;
and that weakly developed parts will in the same manner tend to become
still weaker. Amphigonic reproduction therefore ensures that every
character which is subject to individual fluctuation must appear in
many individuals with a strengthened degree of development, in many
others with a development which is less than normal, while in a still
larger number of individuals the average development will be reached.
Such differences afford the material by means of which natural
selection is able to increase or weaken each character according to the
needs of the species. By the removal of the less well-adapted
individuals, natural selection increases the chance of beneficial
cross-breeding in the subsequent generations.
Every one must admit that, if a species came into existence having only
a small number of individual differences which appeared in the
different parts of different individuals, the number of differences
would increase with each sexually produced generation, until all the
parts in which the variations occurred had received a peculiar
character in all individuals.
Moreover sexual reproduction not only adds to the number of existing
differences, but it also brings them into new combinations, and this
latter consequence is as important as the former.
The former consequence can hardly make itself felt in any existing
species, because in them every part already possesses its peculiar
character in all individuals. The second consequence is, however, more
important, viz. the production of new combinations of individual
characters by sexual reproduction; for, as Darwin has already pointed
out, we must imagine that not only are single characters changed in the
process of breeding, but that probably several, and perhaps very many
characters, are simultaneously modified. No two species, however nearly
allied, differ from each other in but a single character. Even our
eyesight, which has by no means reached the highest pitch of
development, can always detect several, and often very many points of
difference; and if we possessed the powers necessary for making an
absolutely accurate comparison, we should probably find that everything
is different in two nearly allied species.
It is true that a great number of these differences depend upon
correlation, but others must depend upon simultaneous primary changes.
Public-domain text, read in full here on John Shaqi.
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