Essays Upon Heredity and Kindred Biological Problems: Authorised TranslationWeismann, August
Science
Essays Upon Heredity and Kindred Biological Problems: Authorised Translation
Weismann, August
Evolution (Biology); Genetics; Reproduction
Rejuvenescence can be rendered intelligible in theory by some such
metaphor, but considerable difficulties are encountered in the rigid
application of the metaphor to the facts of the case. In the first
place, how is it possible that the motive force can be exhausted by
continual division, while one of its components is being formed afresh
in the same body and during the same time? When thoroughly examined the
loss of the power of division is seen to follow from the loss of the
powers of assimilation, nutrition, and growth. How is it possible that
such a power can be weakened and finally entirely lost while one of its
components is accumulated?
I believe that, instead of accepting such daring assumptions, it is
better to be satisfied with the simple conception of living matter
possessing as attributes the powers of unlimited assimilation and
capacity for reproduction. With such a theory the mere form of
reproduction, whether sexual or asexual, will have no influence upon
the duration of the capacity: for force and matter undergo simultaneous
increase, and are inseparably connected in this as in all other
instances. This theory does not, however, exclude the possible
occurrence of circumstances under which such an association is no
longer necessary.
I could only consent to adopt the hypothesis of rejuvenescence, if it
were rendered absolutely certain that reproduction by division could
never under any circumstances persist indefinitely. But this cannot be
proved with any greater certainty than the converse proposition, and
hence, as far as direct proof is concerned, the facts are equally
uncertain on both sides. The hypothesis of rejuvenescence is, however,
opposed by the fact of parthenogenesis; for if fertilization possesses
in any way the meaning of rejuvenescence, and depends upon the union of
two different forms of force and of matter, which thus produce the
power of reproduction, it follows that we cannot understand how it
happens that the same power of reproduction may be sometimes produced
from one form of matter, alone and unaided. Logically speaking,
parthenogenesis should be as impossible as that either nitric acid or
glycerine should separately produce the effect of nitro-glycerine. The
supposition has indeed been made that in the case of parthenogenesis,
one fertilization is sufficient for a whole series of generations, but
this supposition is not only incapable of proof, but it is contradicted
by the fact that certain eggs which may develope parthenogenetically
are also capable of fertilization. If, in this case, the power of
reproduction were sufficient for development, how is it that the egg is
also capable of fertilization; and if the power were insufficient, how
is it that the egg can develope parthenogenetically? And yet one and
the same egg (in the bee) can develope into a new individual, with or
without fertilization. We cannot escape this dilemma by making the
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