Essays Upon Heredity and Kindred Biological Problems: Authorised TranslationWeismann, August
Science
Essays Upon Heredity and Kindred Biological Problems: Authorised Translation
Weismann, August
Evolution (Biology); Genetics; Reproduction
No objection can be raised against these statements, but Nägeli goes on
to say:—‘But how can such a process of development be explained by the
theory of natural selection, for at each stage in the process the
adaptation was invariably complete. The tube of the corolla and the
“tongue” must have reached, for instance, at a certain time, a length
of 5 or 10 mm. If now the tube of the corolla became longer in some
plants, such an alteration would have been disadvantageous because the
insects would be no longer able to obtain food from them, and would
therefore visit flowers with shorter tubes. Hence, according to the
theory of natural selection, the longer tubes ought to have
disappeared. If on the other hand the “tongue” became longer in some
insects, such a change would be superfluous and should have been given
up, according to the same theory, as unnecessary structural waste. The
simultaneous change in the two structures must, according to the theory
of natural selection, be due to the same principle as that by which
Münchhausen pulled himself out of a bog by means of his own pig-tail.’
But, according to the theory of natural selection, the case appears in
a very different light from that in which it is put by Nägeli. The
flower and the insect do not compete for the greater length of their
respective organs: all through the gradual process, the flower is the
first to lengthen its corolla and the butterfly follows. Their relation
is not like that between a certain species of animal and another which
serves as its prey, where each strives to be the quicker, so that the
speed of both is increased to the greatest possible extent in the
course of generations. Nor do they stand in the same relation as that
obtaining between an insectivorous bird and a certain species of
butterfly which forms its principal food; in such a case two totally
different characters may be continually increased up to their highest
point, e.g. in the butterfly similarity to the dead and fallen leaves
among which it seeks protection when pursued, in the bird keenness of
sight. As long as the latter quality is still capable of increase, so
long will it still be advantageous to any individual butterfly to
resemble the leaf a little more completely than other individuals of
the same species; for it will thus be capable of escaping those birds
which possess a rather keener sight than others. On the other hand, a
bird with rather keener sight will have the greatest chance of catching
the better protected butterflies. It is only in this way that we can
explain the constant production of such extraordinary similarities
between insects and leaves or other parts of plants. At every stage of
growth both the insect and its pursuer are completely adapted to each
other; i.e. they are so far protected and so far successful
respectively, as is necessary to prevent that gradual decrease in the
average number of individuals which would lead to the extermination of
the species[211].
Public-domain text, read in full here on John Shaqi.
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