That which I just now called the fundamental problem of palæontology, the question which has to be {58} settled before any other can be profitably discussed, is this, What is the nature of fossils? Are they, as the healthy common sense of the ancient Greeks appears to have led them to assume without hesitation, the remains of animals and plants? Or are they, as was so generally maintained in the fifteenth, sixteenth, and seventeenth centuries, mere figured stones, portions of mineral matter which have assumed the forms of leaves and shells and bones, just as those portions of mineral matter which we call crystals take on the form of regular geometrical solids? Or, again, are they, as others thought, the products of the germs of animals and of the seeds of plants which have lost their way, as it were, in the bowels of the earth, and have achieved only an imperfect and abortive development? It is easy to sneer at our ancestors for being disposed to reject the first in favour of one or other of the last two hypotheses; but it is much more profitable to try to discover why they, who were really not one whit less sensible persons than our excellent selves, should have been led to entertain views which strike us as absurd. The belief in what is erroneously called spontaneous generation, that is to say, in the development of living matter out of mineral matter, apart from the agency of pre-existing living matter, as an ordinary occurrence at the present day--which is still held by some of us, was universally accepted as an obvious truth by them. They could point to the arborescent forms assumed by hoar-frost and by sundry metallic minerals as evidence of the existence in nature of a "plastic {59} force" competent to enable inorganic matter to assume the form of organised bodies. Then, as every one who is familiar with fossils knows, they present innumerable gradations, from shells and bones which exactly resemble the recent objects, to masses of mere stone which, however accurately they repeat the outward form of the organic body, have nothing else in common with it; and, thence, to mere traces and faint impressions in the continuous substance of the rock. What we now know to be the results of the chemical changes which take place in the course of fossilisation, by which mineral is substituted for organic substance, might, in the absence of such knowledge, be fairly interpreted as the expression of a process of development in the opposite direction--from the mineral to the organic. Moreover, in an age when it would have seemed the most absurd of paradoxes to suggest that the general level of the sea is constant, while that of the solid land fluctuates up and down through thousands of feet in a secular ground swell, it may well have appeared far less hazardous to conceive that fossils are sports of nature than to accept the necessary alternative, that all the inland regions and highlands, in