EvolutionWatson, James A. S. (James Anderson Scott)
History
Evolution
Watson, James A. S. (James Anderson Scott)
Evolution
The process that we have described is that of
sexual reproduction, and its essential features are the same as in
Volvox throughout the whole animal kingdom. The small free-swimming
cells are the male reproductive bodies or sperms, the others are the
female or egg cells. The union of the two produces the fertilised egg,
and the process of union is termed fertilisation. In Volvox, the male
and female elements are sometimes produced by the same individual, at
other times by different ones. Separation of the sexes is no necessary
accompaniment of the process of sexual reproduction, and indeed it is
only in the higher groups of animals that separate sexes are the rule.
The various conditions in Volvox are illustrated in Figs. 28, 29, and
30.
[Illustration: FIG. 30.--Volvox. Portion of a hermaphrodite individual,
showing egg cells (_O, O_{1}_), and sperms (_S_{1} S_{2} S_{3}_).]
The next great groups of animals are, on the one hand, that of the
sponges, and, on the other, that which includes the sea-anemones,
jelly-fishes, corals, etc. At first sight their structure seems vastly
different to that of the Volvox, from some form similar to which they
have probably been derived. The evidence obtained from the study of
their individual development, however, strongly suggests a process by
which we suppose that they evolved from Volvox-like ancestors. We shall
therefore briefly describe the earlier stages of the development of a
coral. The sexually produced individual starts life as a single cell,
the fertilised egg. This divides and redivides until a hollow ball of
cells is produced, which cells, like those of the Volvox, bear cilia.
Although simply spherical in shape, the creature moves by rotating
round a definite axis, like a planet. Moreover, nutriment is absorbed
not by any or every part of the surface, but only by a small area
round the lower pole. Now as development proceeds, the cells at this
pole divide more rapidly than the rest, with the natural result that
the ball begins to get out of shape. The distended portion, however,
develops to the inside, so that one part of the sphere is, as it were,
pushed into the other. When this process has been completed, the
original internal cavity is almost entirely eliminated, and a form is
produced which resembles a double-walled flask or vase. Such a form may
be taken as the fundamental architectural type of the groups that we
are now to consider. The meaning of this further step of Evolution is
again specialisation. The inner layer of cells takes on the functions
of digestion and absorption of food, there having been evolved, in
fact, the simplest possible form of mouth and stomach. Such other
functions as those of locomotion, protection, and support are exercised
by the outer layer. This process is illustrated in Fig. 31.
[Illustration: FIG. 31.--Process of gastrulation in a coral.
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