EvolutionWatson, James A. S. (James Anderson Scott)
History
Evolution
Watson, James A. S. (James Anderson Scott)
Evolution
If now we make a brief general survey of the group to which the Hydra
belongs, we find in it two somewhat strikingly different types. On
the one hand are sedentary forms that resemble, in a general way, the
Hydra; that consist of a tube-shaped body, with the mouth, surrounded
by a ring of tentacles, at the upper end. The sea-anemones and
corals are examples of this type, in which, however, the structure
shows various complexities as compared with that of the Hydra, which
complexities we cannot here pause to describe. On the other hand is the
well-known Medusa form, of which the common jelly-fish is a typical
example. This creature, as is well known, is mushroom shaped, with
tentacles round the edge. The mouth is in the middle of the lower
aspect, at the end of a short 'stalk.' This type is very different in
general appearance from the Hydra or sea-anemone, yet the one form
may be somewhat easily derived from the other; we have only to imagine
that a Hydra is turned upside down, that it is squashed, vertically,
until the internal cavity is greatly reduced, and the circumference,
especially in the region of the tentacles, greatly increased, and we
should have something resembling a Medusa. That the two types are
actually closely related is shown by the fact that there is in the
life-history of one group of Cœlenterates a regular alternation between
the one and the other.
[Illustration: FIG. 35.--Diagram of Medusa.
_rad_, Radial canals, with reproductive bodies, _o_; _r_, ring canal;
_t_, tentacle canal.]
If the above general conception of the structure of the Medusa be
borne in mind, its details will be easily understood. The internal
cavity, instead of being simple, has become complicated, through
the obliteration of certain parts of it, where the upper and lower
walls come in contact. What is left is a comparatively small cavity
immediately above the mouth, a number of symmetrically arranged canals
radiating out from this, and a ring canal connecting the ends of these
with each other. Another special characteristic is that there is a
great mass of gelatinous substance between the outer and inner cell
layers. The reproductive cells, as in the sea-anemones, are produced by
the inner cell layer, and escape by the mouth.
[Illustration: FIG. 36.--Diagrammatic section of Medusa.]
[Illustration: FIG. 37.--Group of Cœlenterates--Medusæ, Sea-anemones,
and Corals.]
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