EvolutionWatson, James A. S. (James Anderson Scott)
History
Evolution
Watson, James A. S. (James Anderson Scott)
Evolution
The worms comprise many greatly divergent groups, and the difference
between the lowest and the highest of these has been produced by many
important steps in Evolution. Of these groups but few immediately
concern us; the first and lowest of those which do, is that of
the Turbellarians, a section of the Platodes or flat-worms. The
Turbellarians are small or microscopic tongue-shaped organisms, of
which the majority of species live on the sea-floor, others however
being found in fresh water. The surface of the body is covered
uniformly with cilia, which serve, in the smaller forms, as organs
of propulsion, while in the larger they appear to have the function
of maintaining a flow of fresh water over the surface, and thus of
assisting respiration. In some respects there has been little advance
from the condition of the Cœlenterate. The digestive cavity is a
simple or more or less divided sac, communicating with the exterior
only by means of the mouth. Unlike the condition of affairs in the
Cœlenterates and Ctenophora, however, the sex glands do not discharge
the reproductive bodies into the digestive cavity, but directly to
the exterior by means of a special opening. Each individual has a pair
of male and a pair of female reproductive glands, but the eggs are
not self-fertilised; nor is the fertilisation of the eggs trusted to
chance and the sea-water, as in the lower groups. Instead there is
a definite exchange of sperms between two individuals, and the eggs
are fertilised before they leave the body. They are also frequently
supplied with a store of nutritive material by a pair of special yolk
glands. A distinct step of progress can thus be recognised in the
arrangements for reproduction. Between the outer skin and the inner
digestive layer is developed a considerable mass of cells, forming
muscular and connective tissue, etc. It will readily be understood that
the development of such thick tissue masses occasions two distinct
new difficulties in the animal economy; for where cells are in direct
contact neither with the digestive layer nor with the exterior, their
nutrition and the removal of their waste products can no longer be
efficiently carried on without special devices. Hence on the one
hand a circulatory system, for the transport of food materials,
and on the other an excretory system, become necessary. The first
of these new departures was not destined to be made until the next
stage of progress; the Turbellarians seem to have temporarily got
over the difficulty, like the Ctenophora, by developing a complex
and ramifying digestive cavity. An excretory system, however, makes
its appearance here. Indeed, the beginnings of such a system can be
seen in the Ctenophora, in which there are small excretory organs
opening into the digestive cavity. The corresponding organs in the
worms, as in all subsequent types, open directly to the outside. In
the Turbellarians these organs, which are termed nephridia, are two in
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