EvolutionWatson, James A. S. (James Anderson Scott)
History
Evolution
Watson, James A. S. (James Anderson Scott)
Evolution
We must now go back and take up the main thread of our story. The next
stage that falls to be described is that of a highly interesting group
of worms known as Enteropneusta, a name signifying 'gut breathers.'
This group contains a very small number of worm forms, which are to
be found burrowing in the sand of the sea floor. A typical example is
the Balanoglossus, a worm of some four inches in length, whose general
appearance is illustrated in Fig. 59. The creature has, as will be
observed, a large muscular snout or proboscis, behind which follows a
small portion called the collar, and behind this again the long body.
The most noteworthy feature of the group, as the name implies, is the
respiratory system. The mouth, which is situated in the region of the
'collar,' leads into a gullet, which is partially divided into an upper
and a lower canal by means of two inwardly projecting longitudinal
folds, one on either side. Only the lower of these canals is used as a
food passage; the upper communicates with the outside by means of
a large number of transverse slits on its sides. Water is continually
being taken in by the mouth and passed along this upper canal, to reach
the outside by way of the gill slits, and in doing so it passes over
the gills, where the blood is circulating in fine capillary vessels.
Here the blood is supplied with oxygen from the water, and is at the
same time relieved of carbonic oxide. This, it will be observed, is
the same method of respiration as that of the fishes. Behind the last
gill slit the digestive canal becomes a simple tube, with two digestive
glands or liver sacs. There are two main blood vessels, the larger
running along above the digestive canal, and the smaller below it, the
two being connected by means of numerous branches. There is a swelling
of the dorsal vessel--a heart--at its forward extremity, in the base
of the proboscis. The nervous system is peculiar; it consists of two
nerve cords, the smaller below the gut and the larger above it--the
latter therefore occupying a position similar to that of the spinal
cord in the vertebrates. Thus in two respects, in its respiratory
and nervous systems, Balanoglossus must be regarded as a highly
extraordinary member of the worm group, and in both its peculiarities
it shows an approach to the vertebrate. There can be little doubt as
to the position of this group as an important connecting link between
the ordinary worms and the vertebrates. Fig. 60 illustrates the main
features of the anatomy.
[Illustration: FIG. 60.--Section of front end of Balanoglossus.
_e_, Snout; _m_, mouth; _h_, heart; _cö_, body cavity; _d_, alimentary
canal; _n_, _n_, nerve cords; _vg_, ventral blood vessel; _dg_, dorsal
blood vessel; _f_, fold dividing the alimentary canal; _vd_, food
canal; _k_, gill slits.]
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account