Second, the polymorphism due to intercrossing is the outcome of
different combinations of hereditary qualities. There are here, De Vries
says, two important classes of facts to be kept strictly
apart,—scientific experiment, and the results of the gardener and of the
cultivator. The experimenter chooses for crossing, species as little
variable as possible; the gardener and cultivator on the other hand
prefer to cross forms of which one at least is variable, because the
variations may be transmitted to the hybrid, and in this way a new form
be produced.
New elementary characters arise in experiments in crossing only through
variability, not through crossing itself.
Third, variability in the ordinary sense, that is, individual
variability, includes those differences between the individual organs
that follow Quetelet’s theory of chance. This kind of variability is
characterized by its presence at all times, in all groups of
individuals.
De Vries recalls Galton’s apt comparison between variability and a
polyhedron which can roll from one face to another. When it comes to
rest on any particular face, it is in stable equilibrium. Small
vibrations or disturbances may make it oscillate, but it returns always
to the same face. These oscillations are like the fluctuating
variations. A greater disturbance may cause the polyhedron to roll over
on to a new face, where it comes to rest again, only showing the ever
present fluctuations around its new centre. The new position corresponds
to a mutation. It may appear from our familiarity with the great changes
that we associate with the idea of discontinuous variability, that a
mutation must also involve a considerable change. Such, however, De
Vries says, is not the case. In fact, numerous mutations are smaller
than the extremes of fluctuating variation. For example, the different
elementary species of _Draba verna_ are less different from each other
than the forms of leaves on a tree. The essential differences between
the two kinds of variation is that the mutation is constant, while the
continuous variation fluctuates back and forth.
The following example is given by De Vries to illustrate the general
point of view in regard to varieties and species. The species _Oxalis
corniculata_ is a “collective” species that lives in New Zealand. It has
been described as having seven well-characterized varieties which do not
live together or have intermediate forms. If we knew only this group,
there would be no question that there are seven good species. But in
other countries intermediate forms exist, which exactly bridge over the
differences between the seven New Zealand forms. For this reason all the
forms have been united in a single species.
Public-domain text, read in full here on John Shaqi.
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