Another example is that of the fern, _Lomaria procera_, from New
Zealand, Australia, South Africa, and South America. If the forms from
only one country be considered, they appear to be different species; but
if all the forms from the different parts of the world be taken into
account, they constitute a connected group, and are united into one
large species.
It will be seen, therefore, that the limits of a collective species are
determined solely by the deficiencies in the genealogical tree of the
elementary species. If all the elementary species in one country were
destroyed, then the forms living in other countries that had been
previously held together because of those which have now been destroyed,
would, after the destruction, become true species. In other words: “The
Linnæan species are formed by the disappearance of other elementary
species, which at first connected all forms. This mode of origin is a
purely historical process, and can never become the subject of
experimental investigation.” Spencer’s famous expression, the “survival
of the fittest,” is incomplete, and should read the “survival of the
fittest species.” It is, therefore, not the study of Linnæan species
that has a physiological interest, but it is the study of the elementary
species of which the Linnæan species are made up, that furnishes the
all-important problem for experimental study.
De Vries gives a critical analysis of a number of cases in which new
races have been formed under domestication. He shows very convincingly
that, whenever the result has been the outcome of the selection of
fluctuating variations, the product that is formed can only be kept to
its highest point of development by the most rigid and ever watchful
care. If selection ceases for only a few generations, the new form sinks
back at once to its original level. Many of our cultivated plants have
really arisen, not by selection of this sort, but by mutations; and
there are a number of recorded cases where the first and sudden
appearance of a new form has been observed. In such cases as these there
is no need for selection, for if left to themselves there is no return
to the original form. If, however, after a new mutation has appeared in
this way, we subject its fluctuating variations to selection, we can
keep the new form up to its most extreme limit, but can do nothing more.
Another means, frequently employed, by which new varieties have been
formed is by bringing together different elementary species under
cultivation. For instance, there are a large number of wild elementary
species of apples, and De Vries believes that our different races of
apples owe their origin in part to these different wild forms. Crossing,
cultivation, and selection have done the rest.
Public-domain text, read in full here on John Shaqi.
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