An interesting case is that of _Daphnia rectirostris_, a small
crustacean living sometimes in fresh water, at other times in water
containing salt and also in salt lakes. There are two forms,
corresponding to the conditions under which they live, and it is said
that the differences are of a kind that suffice to separate species from
each other. In another crustacean, _Branchipus ferox_, the form differs
in a number of points, according to whether it lives in salt or in fresh
water. Schmankewitsch says that, had he not found all transitional
forms, and observed the transformation in cultures, he would have
regarded the two forms as separate species. The oft-quoted case of
Artemia furnishes a very striking example of the influence of the
environment. _Artemia salina_ lives in water whose concentration varies
between 5 and 12 degrees of saltness. When the amount of salt is
increased to 12 degrees, the animal shows certain characteristics like
those of _Artemia milhausenii_, which may live in water having 24 to 25
degrees of saltness. The form _A. salina_ may be further completely
changed into that of _A. milhausenii_ by increasing the amount of salt
to the latter amount.
Among domesticated animals and plants—a few instances of which have been
already referred to—we find a large number of cases in which a change in
the environment produces definite changes in the organism. Darwin has
made a most valuable collection of facts of this kind in his “Animals
and Plants under Domestication.” He believes that domesticated forms are
much more variable than wild ones, and that this is due, in part, to
their being protected from competition, and to their having been removed
from their natural conditions and even from their native country. “In
conformity with this, all our domesticated productions without exception
vary far more than natural species. The hive-bee, which feeds itself,
and follows in most respects its natural habits of life, is the least
variable of all domesticated animals.... Hardly a single plant can be
named, which has long been cultivated and propagated by seed, that is
not highly variable.” “Bud-variation ... shows us that variability may
be quite independent of seminal reproduction, and likewise of reversion
to long-lost ancestral characters. No one will maintain that the sudden
appearance of a moss-rose on a Provence rose is a return to a former
state, ... nor can the appearance of nectarines on peach trees be
accounted for on the principle of reversion.” It is said that
bud-variations are also much more frequent on cultivated than on wild
plants.
Darwin adds: “These general considerations alone render it probable that
variability of every kind is directly or indirectly caused by changed
conditions of life. Or to put the case under another point of view, if
it were possible to expose all the individuals of a species during many
generations to absolutely uniform conditions of life, there would be no
variability.”
Public-domain text, read in full here on John Shaqi.
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