In some cases it has been observed that, in passing from one part of a
continent to another, many or all of the forms of the same group and
even of different groups change in the same way. Allen’s account of the
variations in North American birds and mammals furnishes a number of
striking examples of this kind of change. He finds that, as a rule, the
birds and mammals of North America increase in size from the south
northward. This is true, not only for the individuals of the same
species, but generally the largest species of each genus are in the
north. There are some exceptions, however, in which the increase in size
is in the opposite direction. The explanation of this is that the
largest individuals are almost invariably found in the region where the
group to which the species belongs receives its greatest numerical
development. This Allen interprets as the hypothetical “centre of
distribution of the species,” which is in most cases doubtless also its
original centre of dispersal. If the species has arisen in the north,
then the northern forms are the largest; but if it arose in the south,
the reverse is the case. Thus, most of the species of North America that
live north of Mexico are supposed to have had a northern origin, as
shown by the circumpolar distribution of some of them and by the
relationship of others to Old World species; and in these the largest
individuals of the species of a genus are northern. Conversely, in the
exceptional cases of increase in size toward the south, it can be shown
that the forms have probably had a southern origin.
The Canidæ (wolves and foxes) have their largest representatives, the
world over, in the north. “In North America the family is represented by
six species, the smallest of which (speaking generally) are southern and
the largest northern.” The three species that have the widest ranges
(the gray wolf, the common fox, and the gray fox) show the most marked
differences in size. The skull, for instance, of “the common wolf is
fully one-fifth larger in the northern parts of British America and
Alaska than it is in northern Mexico, where it finds the southern limit
of its habitat. Between the largest northern skull and the largest
southern skull there is a difference of about thirty-five per cent of
the mean size. Specimens from the intermediate region show a gradual
intergradation between the extremes, although many of the examples from
the upper Missouri country are nearly as large as those from the extreme
north.” The common fox is about one-tenth larger, on the average, in
Alaska than it is in New England. The gray fox, whose habitat extends
from Pennsylvania southward to Yucatan, has an average length of skull
of about five inches in the north, and less than four in Central
America—about ten per cent difference.
Public-domain text, read in full here on John Shaqi.
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