A radial type of structure is often found in fixed forms, and in some
floating forms, like the jellyfish. In a fixed form, a sea-anemone, for
instance, the conditions around the free end and the fixed end of the
body are entirely different, and we find that these two ends are also
different. The free end contains the special sense-organs, the mouth,
tentacles, etc.; while the fixed end contains the organ for attachment.
It is evident that the free end is exposed to the same conditions in all
directions, and it may seem probable that this will account for the
radial symmetry of the anemone. There are also a few free forms, the
sea-urchin for instance, that have a radial symmetry. Whether their
ancestors were fixed forms, for which there is some evidence, we do not
know definitely; but, even if this is true, it does not affect the main
point, namely, that, although at present free to move, the sea-urchin is
radially symmetrical. But when we examine its method of locomotion, we
find that it moves indifferently in any direction over a solid surface;
that is, it keeps its oral face against a solid object, and moves over
the surface in any direction. Under these circumstances the same
external conditions will act equally upon all sides of the body. In
contrast to these common sea-urchins, there are two other related
groups, in which, although traces of a well-marked radial symmetry are
found, the external form has been so changed that a secondary bilateral
form has been superimposed on it. These are the groups of the
clypeasters and the spatangoids, and it is generally supposed that their
forefathers were radially symmetrical forms like the ordinary forms of
sea-urchins. These bilateral forms move in the direction of their plane
of symmetry, but we have no means of knowing whether they first became
bilateral and, in consequence, now move in the direction of the median
plane, or whether they acquired the habit of moving in one direction,
and in consequence acquired a bilateral symmetry. It seems more probable
that the form changed first, for otherwise it is difficult to see why a
change of movement in one direction should ever have taken place.
Public-domain text, read in full here on John Shaqi.
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