The radially symmetrical form is characteristic of many flowers that
stand on the ends of their stalks. They also will be subjected to
similar external influences in all directions. Many flowers, on the
other hand, are bilaterally symmetrical. Some of these forms are of such
a sort that they are generally interpreted as having been acquired in
connection with the visits of insects. Be this as it may, it is still
not clear why, if the flowers are terminal, insects should not approach
them equally from every direction. If the flowers are not terminal, as,
in fact, many of them are not, their relation to the surroundings is
bilateral with respect to internal as well as to external conditions.
The former, rather than the latter, may have produced the bilateral form
of the flower. Here also we meet with the problem as to whether the
flowers, being lateral in position, have assumed a bilateral form
because their internal relations were bilateral; or whether an external
relation, for example, the visits of insects, has been the principle
cause of their becoming bilateral.
[Illustration:
Fig. 4.—A, right and left claws of lobster;
B, of the fiddler-crab; and
C, of Alpheus.]
In some bilateral forms the right and left sides may be unsymmetrical in
certain organs. Right and left handedness in man is the most familiar
example, although the structural difference on which this rests is not
very obvious. More striking is the difference in the two big claws of
the lobster (Fig. 4 A). One of the two claws is flat and has a fine
saw-toothed edge. The other is thicker and has rounded knobs instead of
teeth. It is said that these two claws are used by the lobster for
different purposes,—the heavy one for crushing and for holding on, and
the narrower for cutting up the food. If this is true, then we find a
symmetrical organism becoming unsymmetrical, and in consequence it takes
advantage of its asymmetry by using its right and left claws for
different purposes.
More striking still is the difference in the size of the right and left
claws in a related form, Alpheus—a crayfish-like form that lives in the
sea. With the larger claw (Fig. 4 C) it makes a clicking sound that can
be heard for a long distance. In some of the crabs the difference in the
size of the two claws is enormous, as in the male fiddler-crab, for
example (Fig. 4 B). One of the claws is so big and unwieldy that it must
put the animal at a distinct disadvantage. Its use is unknown, although
it has been suggested that it is a secondary sexual character.
Public-domain text, read in full here on John Shaqi.
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