Evolution and Classification of the Pocket Gophers of the Subfamily Geomyinae — John Shaqi
Evolution and Classification of the Pocket Gophers of the Subfamily GeomyinaeRussell, Robert J.
Science
Evolution and Classification of the Pocket Gophers of the Subfamily Geomyinae
Russell, Robert J.
Pocket gophers
_Pliogeomys buisi_ Hibbard (1954:353) was found in the Buis Ranch
local fauna, of latest middle Pliocene, on the west side of Buckshot
Arroyo, Beaver County, Oklahoma. The original material included a
right ramus bearing the premolar and first two molars (the holotype)
and five isolated premolars and molars. One of the molars is slightly
worn and from an immature individual. One premolar is a deciduous
tooth. Hibbard (_op. cit._:342) identified the beds from which he
obtained the Buis Ranch local fauna as from the lowermost part of the
Upper Pliocene. Moreover, he judged the Buis Ranch local fauna to be
only slightly older than the Saw Rock Canyon local fauna of Seward
County in southwestern Kansas. Previously (Hibbard, 1953:408-410),
the Saw Rock Canyon local fauna had been assessed as older than
the Rexroad local faunas (latest late Pliocene) and, therefore,
representative of the early part of the late Pliocene. More recently,
Hibbard (1956:164) identified the Buis Ranch beds as part of the
Ogallala Formation, which here occurs unconformably just beneath the
Rexroad Formation (composed of strata nearly all of late Pliocene
age). Therefore, he regarded the Buis Ranch beds as latest middle
Pliocene in age. Hibbard (1954:356) suggested that pocket gopher
remains from the Saw Rock Canyon local fauna were referable to
_Pliogeomys buisi_, and, in effect, tentatively assigned them to
_Pliogeomys_ (in his description of the genus Hibbard remarked that
the upper incisor is bisulcate as in _Geomys_, and the only upper
incisor that he mentions was one of the Saw Rock Canyon fossils
and not part of the Buis Ranch material). _Pliogeomys_ has closer
affinities with modern pocket gophers of the tribe Geomyini than it
does with the middle Pliocene genus _Pliosaccomys_.
The pocket gopher fauna known from the late Pliocene was more varied
than the faunas known from any earlier time. In addition to the
extinct _Pliogeomys_, which occurs in early late Pliocene (see
discussion above), the living genera _Zygogeomys_, _Geomys_,
_Pappogeomys_ (in the sense used on p. 534), and _Thomomys_
first appear in the late Pliocene. The only other living genus,
_Orthogeomys_, makes its first appearance in the late Pleistocene.
The earliest record of the genus _Thomomys_ is based on a fragment of
a left mandibular ramus bearing p4 and m1, _Thomomys gidleyi_ Wilson
(1933b:122), from the Hagerman local fauna of Twin Falls County,
Idaho. Wilson (_loc. cit._) was uncertain as to age (late Pliocene or
early Pleistocene) but subsequently (1937:38 and 67-70) settled on the
middle part of the late Pliocene. Hibbard (1958:11) later considered
the age as early Pleistocene (suggesting that the deposits accumulated
in the Aftonian interglacial interval) but subsequently (Hibbard _et
al._, 1965:512), on the basis of potassium argon age determinations,
also settled on late Pliocene.
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