Evolution and Classification of the Pocket Gophers of the Subfamily Geomyinae — John Shaqi
Evolution and Classification of the Pocket Gophers of the Subfamily GeomyinaeRussell, Robert J.
Science
Evolution and Classification of the Pocket Gophers of the Subfamily Geomyinae
Russell, Robert J.
Pocket gophers
Gill, in 1872 (p. 71), proposed a classification essentially like
Baird's of 1858, but Gill raised Baird's subfamilies to the rank
of family (see Table 1). In referring all pocket gophers to the
Geomyidae, Gill used that name as a family term for the first time.
Also he established the superfamily Saccomyoidea to include his two
families, Geomyidae and Saccomyidae; therefore, the Saccomyoidea was
equivalent to the group Saccomyina of Waterhouse (1848) and the
Saccomyidae of Baird (1858). Coues (1877), in his classic monograph of
the Geomyidae followed the arrangement proposed by Gill in treating
the pocket gophers as a family. Alston in 1876 proposed another
classification based on Baird (1858), with two subfamilies, the
Geomyinae and the Heteromyinae, united together in the family
Geomyidae; thus, he recognized that the genus _Saccomys_ Frédéric
Cuvier, 1823, was a synonym of _Heteromys_ Desmarest, 1817, as had
been pointed out by Gray (1868:201) and Peters (1874:356). Coues
(1877:487-490) acknowledged the invalidity of the genus _Saccomys_,
but refused to give up the name in supergeneric classification. Winge,
first in 1887 and subsequently in 1924, classified the geomyids and
heteromyids together in the family Saccomyidae as did Baird in 1858,
and like Coues, Winge too ignored the synonymy of _Saccomys_ with
_Heteromys_ and insisted on retaining the technical terms Saccomyidae
and Saccomyini.
Up to the time of Merriam's classic revision of the Recent Geomyidae
in 1895 all the known species of living pocket gophers were referred
to two genera, _Geomys_ and _Thomomys_. Merriam described much new
material, especially from México and Central America, and proposed
seven new genera (see Table 1). His complete and detailed study of the
dentitions and osteology of the skull remains today as the definitive
work on this subject, and is the point where most studies of the
Geomyidae must begin. His treatment of the Recent genera survived for
52 years without change until Hooper (1946:397) arranged _Platygeomys_
as a synonym of _Cratogeomys_. However, Merriam's genera have been
recognized in all subsequent classifications except for the current
review (see Table 1).
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