Evolution and Classification of the Pocket Gophers of the Subfamily GeomyinaeRussell, Robert J.
Science
Evolution and Classification of the Pocket Gophers of the Subfamily Geomyinae
Russell, Robert J.
Pocket gophers
B Anterior margin of mesopterygoid fossa even with plane of
posterior wall of M3; postorbital bar weakly developed;
anteroposterior occlusal length of M3 equal to, or less
than, combined length of M1 and M2.
Subgenus _Heterogeomys_ p. 530
B´ Anterior margin of mesopterygoid fossa decidedly behind
plane of posterior wall of M3; postorbital bar strongly
developed; anteroposterior occlusal length of M3 more
than combined length of M1 and M2.
Subgenus _Macrogeomys_ p. 531
Subgenus =Orthogeomys= Merriam
1895. _Orthogeomys_ Merriam, N. Amer. Fauna, 8:172, January 31.
_Type._--_Geomys scalops_ Thomas, 1894, from Tehuantepec, Oaxaca,
México.
_Chronologic range._--Known only from the Recent.
_Description._--Skull elongated and narrow (many skulls of nearly
uniform breadth throughout), being extreme in dolichocephalic
specializations; mandibles long and narrow, rami not spreading
laterally, being more nearly parallel-sided than in other subgenera;
angular processes short; breadth across zygomata not significantly
exceeding breadth across mastoid processes (in many skulls
considerably less); interorbital area remarkably broad, lacking deep
constriction; frontals between orbits greatly inflated laterally,
postorbital prominence inconspicuous; mesopterygoid fossa extending to
level of posterior margin of M3; I having sulcus broader than in other
subgenera, mostly on inner half of anterior surface but sometimes
overlapping mid-line; enamel plate lacking from posterior wall of P4,
rarely retaining narrow vestige near lingual border of posterior loph;
M3 having distinct heel, bicolumnar pattern with inner re-entrant fold
usually minute, occlusal length less than in other subgenera, length
less than combined lengths of M1-2; hair generally coarse, sometimes
hispid, sparse, in lowland forms, so sparse as to impart appearance of
nakedness.
_Referred species and subspecies._--Fourteen taxa:
_Orthogeomys grandis alleni_ Nelson and Goldman, 1930. Jour. Mamm.,
11:156, May 9. Type from near Acapulco, 2000 ft., Guerrero.
_Orthogeomys grandis annexus_ Nelson and Goldman, 1933. Proc. Biol.
Soc. Washington, 46:195, October 26. Type from Tuxtla
Gutierrez, 2600 ft., Chiapas.
_Orthogeomys grandis carbo_ Goodwin, 1956. Amer. Mus. Novit.,
1757:5, March 8. Type from Excurano, 2500 ft., Cerro de San
Pedro, 20 km. W Mixtequilla, Oaxaca.
_Orthogeomys grandis felipensis_ Nelson and Goldman, 1930. Jour.
Mamm., 11:157, May 9. Type from Cerro San Felipe, 10 mi. N
Oaxaca, Oaxaca.
_Orthogeomys grandis huixtlae_ Villa, 1944. Anal. Inst. Biol. Univ.
Nac. México, 15:319. Type from Finca Lubeca, 12 km. NE
Huixtla, 850 m., Chiapas.
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