Evolution and Classification of the Pocket Gophers of the Subfamily GeomyinaeRussell, Robert J.
Science
Evolution and Classification of the Pocket Gophers of the Subfamily Geomyinae
Russell, Robert J.
Pocket gophers
_Orthogeomys grandis grandis_ (Thomas, 1893). Ann. Mag. Nat. Hist.,
ser. 6, 12:270, October. Type from Dueñas, Guatemala.
_Orthogeomys grandis latifrons_ Merriam, 1895. N. Amer. Fauna,
8:178, January 31. Type from Guatemala, exact locality
unknown.
_Orthogeomys grandis nelsoni_ Merriam, 1895. N. Amer. Fauna, 8:176,
January 31. Type from Mt. Zempoaltepec, 8000 ft., Oaxaca.
_Orthogeomys grandis pluto_ Lawrence, 1933. Proc. New England Zool.
Club, 13:66, May 8. Type from Cerro Cantoral, north of
Tegucigalpa, Honduras.
_Orthogeomys grandis scalops_ (Thomas, 1894). Ann. Mag. Nat. Hist.,
ser. 6, 13:437, May. Type from Tehuantepec, Oaxaca.
_Orthogeomys grandis soconuscensis_ Villa, 1949. Anal. Inst. Biol.
Univ. Nac. México, 19:267, April 8. Type from Finca
Experanza, 710 m., 45 km. (by road) NW Huixtla, Chiapas.
_Orthogeomys grandis guerrerensis_ Nelson and Goldman, 1930. Jour.
Mamm., 11:158, May 9. Type from El Limón, in valley of Río de
las Balsas approximately 20 mi. NW La Unión, Guerrero.
_Orthogeomys cuniculus_ Elliot, 1905. Proc. Biol. Soc. Washington,
18:234, December 9. Type from Zanatepec, Oaxaca.
_Orthogeomys pygacanthus_ Dickey, 1928. Proc. Biol. Soc. Washington,
41:9, February 1. Type from Cacaguatique, 3500 ft., Dept. San
Miguel, El Salvador.
Subgenus =Heterogeomys= Merriam
1895. _Heterogeomys_ Merriam, N. Amer. Fauna, 8:179, January 21.
_Type._--_Geomys hispidus_ Le Conte, 1852, from near Jalapa, Veracruz.
_Chronologic range._--Late Pleistocene, Wisconsin deposits (San
Josecito Cave local fauna, Nuevo León) to the Recent.
_Description._--Skull dolichocephalic (less so than in the other
subgenera); zygomata more widely spreading than in _Orthogeomys_;
ramus and angular process short; interorbital area noticeably
constricted; frontals between orbits neither exceptionally broad or
inflated; mesopterygoid fossa extending to level of posterior margin
of M3; I having sulcus on inner third of anterior surface usually
narrower than in subgenus _Orthogeomys_; enamel plate on posterior
wall of P4 restricted to lingual half of loph; M3 distinctly
biprismatic, posterior loph usually circumscribed by shallow inner
re-entrant fold and outer deep fold well developed in all members of
genus; posterior loph forming conspicuous heel longer than in subgenus
_Orthogeomys_; occlusal length equal to or slightly less than combined
lengths of M1-2; hair coarse and hispid but never so sparse as to
impart appearance of nakedness.
_Referred species and subspecies._--Eleven taxa:
*_Orthogeomys onerosus_ (Russell, 1960). Univ. Kansas Publ., Mus.
Nat. Hist., 9 (21):544, January 14. Type from San Josecito Cave
local fauna, Upper Pleistocene, Nuevo León.
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