If we include under the expression of the emotions not only the
premonitory symptoms of the initial phases of the organic and mental
state, not only the signs or conditions of half-tide emotion, but the
full-tide manifestation of an emotion which dominates the situation,
we are naturally led on to the consideration of many of the phenomena
which are discussed under the head of sexual selection. The subject is
difficult and complex, and it was treated by Darwin with all the
strength he could summon to the task. It can only be dealt with here
from a special point of view--that which may serve to illustrate the
influence of certain mental factors on the course of evolution. From
this point of view too much stress can scarcely be laid on the
dominance of emotion during the period of courtship and pairing in the
more highly organised animals. It is a period of maximum vigour,
maximum activity, and, correlated with special modes of behaviour and
special organic and visceral accompaniments, a period also of maximum
emotional excitement. The combats of males, their dances and aerial
evolutions, their elaborate behaviour and display, or the flood of
song in birds, are emotional expressions which are at any rate
coincident in time with sexual periodicity. From the combat of the
males there follows on Darwin's principles the elimination of those
which are deficient in bodily vigour, deficient in special structures,
offensive or protective, which contribute to success, deficient in the
emotional supplement of which persistent and whole-hearted fighting is
the expression, and deficient in alertness and skill which are the
outcome of the psychological development of the powers of perception.
Few biologists question that we have here a mode of selection of much
importance, though its influence on psychological evolution often
fails to receive its due emphasis. Mr. Wallace[174] regards it as "a
form of natural selection"; "to it," he says, "we must impute the
development of the exceptional strength, size, and activity of the
male, together with the possession of special offensive and defensive
weapons, and of all other characters which arise from the development
of these or are correlated with them." So far there is little
disagreement among the followers of Darwin--for Mr. Wallace, with fine
magnanimity, has always preferred to be ranked as such,
notwithstanding his right, on which a smaller man would have
constantly insisted, to the claim of independent originator of the
doctrine of natural selection. So far with regard to sexual selection
Darwin and Mr. Wallace are agreed; so far and no farther. For Darwin,
says Mr. Wallace,[175] "has extended the principle into a totally
different field of action, which has none of that character of
constancy and of inevitable result that attaches to natural selection,
including male rivalry; for by far the larger portion of the
phenomena, which he endeavours to explain by the direct action of
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