sexual selection, can only be so explained on the hypothesis that the immediate agency is female choice or preference. It is to this that he imputes the origin of all secondary sexual characters other than weapons of offence and defence.... In this extension of sexual selection to include the action of female choice or preference, and in the attempt to give to that choice such wide-reaching effects, I am unable to follow him more than a very little way." Into the details of Mr. Wallace's criticisms it is impossible to enter here. We cannot discuss either the mode of origin of the variations in structure which have rendered secondary sexual characters possible or the modes of selection other than sexual which have rendered them, within narrow limits, specifically constant. Mendelism and mutation theories may have something to say on the subject when these theories have been more fully correlated with the basal principles of selection. It is noteworthy that Mr. Wallace says:[176] "Besides the acquisition of weapons by the male for the purpose of fighting with other males, there are some other sexual characters which may have been produced by natural selection. Such are the various sounds and odours which are peculiar to the male, and which serve as a call to the female or as an indication of his presence. These are evidently a valuable addition to the means of recognition of the two sexes, and are a further indication, that the pairing season has arrived; and the production, intensification, and differentiation of these sounds and odours are clearly within the power of natural selection. The same remark will apply to the peculiar calls of birds, and even to the singing of the males." Why the same remark should not apply to their colours and adornments is not obvious. What is obvious is that "means of recognition" and "indication that the pairing season has arrived" are dependent on the perceptive powers of the female who recognises and for whom the indication has meaning. The hypothesis of female preference, stripped of the aesthetic surplusage which is psychologically both unnecessary and unproven, is really only different in degree from that which Mr. Wallace admits in principle when he says that it is probable that the female is pleased or excited by the display.