We must briefly refer here to the views--much discussed in the last
decade--of H. de Vries, who believes that the roots of transformation
must be sought for in _saltatory variations arising from internal
causes_, and distinguishes such _mutations_, as he has called them,
from ordinary individual variations, in that they breed true, that is,
with strict in-breeding they are handed on pure to the next
generation. I have elsewhere endeavoured to point out the weaknesses
of this theory,[33] and I am the less inclined to return to it here
that it now appears[34] that the far-reaching conclusions drawn by de
Vries from his observations on the Evening Primrose, _Oenothera
lamarckiana_, rest upon a very insecure foundation. The plant from
which de Vries saw numerous "species"--his "mutations"--arise was not,
as he assumed, a _wild species_ that had been introduced to Europe
from America, but was probably a hybrid form which was first
discovered in the Jardin des Plantes in Paris, and which does not
appear to exist anywhere in America as a wild species.
This gives a severe shock to the "Mutation theory," for the other
_actually wild_ species with which de Vries experimented showed no
"mutations" but yielded only negative results.
Thus we come to the conclusion that Darwin[35] was right in regarding
transformations as taking place by minute steps, which, if useful, are
augmented in the course of innumerable generations, because their
possessors more frequently survive in the struggle for existence.
(_b_) _Selection-value of the initial steps_
Is it possible that the insignificant deviations which we know as
"individual variations" can form the beginning of a process of
selection? Can they decide which is to perish and which to survive? To
use a phrase of Romanes, can they have _selection-value_?
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