This illustration is entirely apt, the only difference between the
two cases consisting in the fact that the variation in the flower is
not a useful, but a disadvantageous one, which can only be preserved
by artificial selection on the part of the gardener, while the
transformations that have taken place parallel with the sterility of
the ants are useful, since they procure for the colony an advantage in
the struggle for existence, and they are therefore preserved by
natural selection. Even the sterility itself in this case is not
disadvantageous, since the fertility of the true females has at the
same time considerably increased. We may therefore regard the sterile
forms of ants, which have gradually been adapted in several directions
to varying functions, _as a certain proof_ that selection really takes
place in the germ-cells of the fathers and mothers of the workers, and
that _special complexes of primordia_ (_ids_) are present in the
workers and in the males and females, and these complexes contain the
primordia of the individual parts (_determinants_). But since all
living entities vary, the determinants must also vary, now in a
favourable, now in an unfavourable direction. If a female produces
eggs, which contain favourably varying determinants in the worker-ids,
then these eggs will give rise to workers modified in the favourable
direction, and if this happens with many females, the colony concerned
will contain a better kind of worker than other colonies.
I digress here in order to give an account of the intimate processes,
which, according to my view, take place within the germ-plasm, and
which I have called "_germinal selection_." These processes are of
importance since they form the roots of variation, which in its turn
is the root of natural selection. I cannot here do more than give a
brief outline of the theory in order to show how the Darwin-Wallace
theory of selection has gained support from it.
With others, I regard the minimal amount of substance which is
contained within the nucleus of the germ-cells, in the form of rods,
bands, or granules, as the _germ-substance_ or _germ-plasm_, and I
call the individual granules _ids_. There is always a multiplicity of
such ids present in the nucleus, either occurring individually, or
united in the form of rods or bands (chromosomes). Each id contains
the primary constituents of a _whole_ individual, so that several ids
are concerned in the development of a new individual.
Public-domain text, read in full here on John Shaqi.
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