One of the chief preliminary postulates of sexual selection is the
unequal number of individuals in the two sexes, for if every male
immediately finds his mate there can be no competition for the
possession of the female. Darwin has shown that, for the most part,
the inequality between the sexes is due simply to the fact that there
are more males than females, and therefore the males must take some
pains to secure a mate. But the inequality does not always depend on
the numerical preponderance of the males, it is often due to polygamy;
for, if one male claims several females, the number of females in
proportion to the rest of the males will be reduced. Since it is
almost always the males that are the wooers, we must expect to find
the occurrence of secondary sexual characters chiefly among them, and
to find it especially frequent in polygamous species. And this is
actually the case.
If we were to try to guess--without knowing the facts--what means the
male animals make use of to overcome their rivals in the struggle for
the possession of the female, we might name many kinds of means, but
it would be difficult to suggest any which is not actually employed in
some animal group of other. I begin with the mere difference in
strength, through which the male of many animals is so sharply
distinguished from the female, as, for instance, the lion, walrus,
"sea-elephant," and others. Among these the males fight violently for
the possession of the female, who falls to the victor in the combat.
In this simple case no one can doubt the operation of selection, and
there is just as little room for doubt as to the selection-value of
the initial stages of the variation. Differences in bodily strength
are apparent even among human beings, although in their case the
struggle for the possession of the female is no longer decided by
bodily strength alone.
Combats between male animals are often violent and obstinate, and the
employment of the natural weapons of the species in this way has led
to perfecting of these, e.g. the tusks of the boar, the antlers of the
stag, and the enormous, antler-like jaws of the stag-beetle. Here
again it is impossible to doubt that variations in these organs
presented themselves, and that these were considerable enough to be
decisive in combat, and so to lead to the improvement of the weapon.
Public-domain text, read in full here on John Shaqi.
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