We come now to the _excitants_; that is, to the group of sexual
characters whose origin through processes of selection has been most
frequently called in question. We may cite the _love-calls_ produced
by many male insects, such as crickets and cicadas. These could only
have arisen in animal groups in which the female did not rapidly flee
from the male, but was inclined to accept his wooing from the first.
Thus, notes like the chirping of the male cricket serve to entice the
females. At first they were merely the signal which showed the
presence of a male in the neighbourhood, and the female was gradually
enticed nearer and nearer by the continued chirping. The male that
could make himself heard to the greatest distance would obtain the
largest following, and would transmit the beginnings, and, later, the
improvement of his voice to the greatest number of descendants. But
sexual excitement in the female became associated with the hearing of
the love-call, and then the sound-producing organ of the male began to
improve, until it attained to the emission of the long-drawn-out soft
notes of the mole-cricket or the maenad-like cry of the cicadas. I
cannot here follow the process of development in detail, but will call
attention to the fact that the original purpose of the voice, the
announcing of the male's presence, became subsidiary, and the exciting
of the female became the chief goal to be aimed at. The loudest
singers awakened the strongest excitement, and the improvement
resulted as a matter of course. I conceive of the origin of bird-song
in a somewhat similar manner, first as a means of enticing, then of
exciting the female.
Public-domain text, read in full here on John Shaqi.
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