In this domain, then, the theory of selection finds nothing but
corroboration, and it would be impossible to substitute for it any
other explanation, which now that the facts are so well known, could
be regarded as a serious rival to it. That selection is a factor, and
a very powerful factor in the evolution of organisms, can no longer be
doubted. Even although we cannot bring forward formal proofs of it _in
detail_, cannot calculate definitely the size of the variations which
present themselves, and their selection-value, cannot, in short,
reduce the whole process to a mathematical formula, yet we must assume
selection, because it is the only possible explanation applicable to
whole classes of phenomena, and because, on the other hand, it is made
up of factors which we know can be proved actually to exist, and
which, _if_ they exist, must of logical necessity coöperate in the
manner required by the theory. _We must accept it because the
phenomena of evolution and adaptation must have a natural basis, and
because it is the only possible explanation of them._[53]
Many people are willing to admit that selection explains adaptations,
but they maintain that only a part of the phenomena are thus
explained, because everything does not depend upon adaptation. They
regard adaptation as, so to speak, a special effort on the part of
Nature, which she keeps in readiness to meet particularly difficult
claims of the external world on organisms. But if we look at the
matter more carefully we shall find that adaptations are by no means
exceptional, but that they are present everywhere in such enormous
numbers, that it would be difficult in regard to any structure
whatever, to prove that adaptation had _not_ played a part in its
evolution.
How often has the senseless objection been urged against selection
that it can create nothing, it can only reject. It is true that it
cannot create either the living substance or the variations of it;
both must be given. But in rejecting one thing it preserves another,
intensifies it, combines it, and in this way _creates_ what is new.
_Everything_ in organisms depends on adaptation; that is to say,
everything must be admitted through the narrow door of selection,
otherwise it can take no part in the building up of the whole. But, it
is asked, what of the direct effect of external conditions,
temperature, nutrition, climate and the like? Undoubtedly these can
give rise to variations, but they too must pass through the door of
selection, and if they cannot do this they are rejected, eliminated
from the constitution of the species.
Public-domain text, read in full here on John Shaqi.
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