Evolution: Its nature, its evidence, and its relation to religious thoughtLeConte, Joseph
Religion
Evolution: Its nature, its evidence, and its relation to religious thought
LeConte, Joseph
Evolution (Biology); Evolution (Biology) -- Religious aspects; Religion and science
Take another example: The broad, flat, elliptical, green masses so
characteristic of the cactus family, and usually called their leaves,
are indeed _analogous_ to leaves in color, form, and function; for
they are green and flat, and assimilate carbonic acid and water (CO2
and H2O) like leaves. But they are not, in truth, leaves, but modified
stems, for they have the essential structure of stems, with their
pith, wood, medullary rays, and bark, and may be traced through all
gradations into the ordinary cylindrical form of stems. Where are
their leaves, then? Their spines are their abortive leaves. These
are arranged spirally like leaves, and bear buds in their axils like
leaves. They are, in truth, leaves, modified to perform the function of
defensive armor; while their function has been delegated to the stem
flattened for this purpose.
[Illustration:
FIG. 3.--A branch of young acacia, showing change from one form of
leaf to the other; _a_, _b_, _c_, _d_, successive stages of
change; _l_, _s_, leaf stalk which gradually changes into the
blade in _c_, _d_, and _e_.
]
One more example: The acacias, of which there are fifteen to twenty
species in California, introduced from Australia, form two groups
having extremely different styles of leaves. We will call them the
feather-leaved and the simple-leaved acacias. In the former, the leaves
are very finely bipinnate, and the general aspect of the foliage is
extremely feathery and graceful. In the latter the leaves are simple,
ovate, and, curiously enough, set on edge; and the general aspect
of the tree is therefore rather stiff. It seems at first incredible
that leaves so different and aspects so diverse should belong to
plants of the same genus. But a little close examination shows that,
as usual, the botanists are right and the popular judgment wrong.
The plumose-leaf is the normal leaf-form for this genus. The simple
leaf is not only abnormal, but in a homological sense is not a
leaf at all--i. e., it does not correspond to the part called the
_blade_ in ordinary simple leaves of other trees. In the seedling of
the simple-leaved acacias, and sometimes for a considerable time in
the young tree, the leaves are all plumose. As the tree matures it
gradually changes its dress and puts on its _toga virilis_. The gradual
change from the one form to the other may easily be traced in the same
tree, and even often in the same branch (Fig. 3). The steps of the
change (_a_, _b_, _c_, and _d_) are shown in the following figure,
drawn from nature. It is seen, by bare inspection of the figure, that
the so-called leaf, _d_, of the simple-leaved acacias, is really the
vertically-expanded leaf-stalk, _l_, _s_, the true leaf or blade being
wholly aborted. The whole structure of this so-called leaf is different
from that of a true blade. For example, its style of ribbing is
parallel, its position is edgewise to the sky, its palisade cells are
on both sides alike, etc.
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