In the Amphipoda, to which the above-mentioned naturalists correctly
refer the Caprellidæ and Cyamidæ (Latreille’s _Læmodipoda_), the heart
has always the same position; it extends in the form of a long tube
through the six segments following the head, and has three pairs of
fissures, furnished with valves, for the entrance of the blood,
situated in the second, third, and fourth of these segments. It was
found to be of this structure by La Valette in _Niphargus_ (_Gammarus
puteanus_), and by Claus in _Phronima_; and I have found it to be the
same in a considerable number of species belonging to the most
different families.[1]
The sole unimportant exception which I have hitherto met with is
presented by the genus _Brachyscelus,_[2] in which the heart possesses
only two pairs of fissures, as it extends forward only into the second
body-segment, and is destitute of the pair of fissures situated in this
segment in other forms.[3]
Considering this uniformity presented by the heart in the entire order
of the Amphipoda, it cannot but seem very remarkable, that in the very
next order of the Isopoda, we find it to be one of the most changeable
organs.
In the cheliferous Isopods (_Tanais_) the heart resembles that of the
Amphipoda in its elongated tubular form, as well as in the number and
position of the fissures, but with this difference, that the two
fissures of each pair do not lie directly opposite each other.
Fig. 14. Heart of a young Cassidina. Fig. 15. Heart of a young
Anilocra. Fig. 16. Abdomen of the male of Entoniscus Cancrorum. h.
Heart. l. Liver. Fig. 14. Heart of a young _Cassidina._ Fig. 15. Heart
of a young _Anilocra._ Fig. 16. Abdomen of the male of _Entoniscus
Cancrorum. h._ Heart. _l._ Liver.
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