[9] I have not mentioned the Pycnogonidæ, because I do not regard them
as Crustacea; nor the Xiphosura and Trilobites, because, having never
investigated them myself, I knew too little about them, and especially
because I am unacquainted with the details of the explanations given
by Barrande of the development of the latter. According to Mr. Spence
Bate “the young of Trilobites are of the Nauplius-form.”)
CHAPTER X.
ON THE PRINCIPLES OF CLASSIFICATION.
Perhaps some one else, more fortunate than myself, may be able, even
without Darwin, to find the guiding clue through the confusion of
developmental forms, now so totally different in the nearest allies,
now so surprisingly similar in members of the most distant groups,
which we have just cursorily reviewed. Perhaps a sharper eye may be
able, with Agassiz, to make out “the plan established from the
beginning by the Creator,”[1] who may have written here, as a
Portuguese proverb says “straight in crooked lines.”[2] I cannot but
think that we can scarcely speak of a general plan, or typical mode of
development of the Crustacea, differentiated according to the separate
Sections, Orders, and Families, when, for example, among the Macrura,
the River Crayfish leaves the egg in its permanent form; the Lobster
with Schizopodal feet; _Palæmon,_ like the Crabs, as a Zoëa; and
_Penéus,_ like the Cirripedes, as a Nauplius,—and when, still, within
this same sub-order Macrura, _Palinurus, Mysis_ and _Euphausia_ again
present different young forms,—when new limbs sometimes sprout forth as
free rudiments on the ventral surface, and are sometimes formed beneath
the skin which passes smoothly over them, and both modes of development
are found in different limbs of the same animal and in the same pair of
limbs in different animals,—when in the Podophthalma the limbs of the
thorax and abdomen make their appearance sometimes simultaneously, or
sometimes the former and sometimes the latter first, and when further
in each of the two groups the pairs sometimes all appear together, and
sometimes one after the other,—when, among the Hyperina, a simple foot
becomes a chela in _Phronima_ and a chela a simple foot in
_Brachyscelus,_ etc.
And yet, according to the teaching of the school, it is precisely in
youth, precisely in the course of development, that the “Type” is
mostly openly displayed. But let us hear what the Old School has to
tell us as to the significance of developmental history, and its
relation to comparative anatomy and systematic zoology.
Let two of its most approved masters speak.
Public-domain text, read in full here on John Shaqi.
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