Better seeds--that is, those that produce stronger and more
fruitful plants--often result when the _pollen comes from another
flower_. Fertilization effected between different flowers is
=cross-fertilization=; that resulting from the application of
pollen to pistils in the same flower is =close-fertilization= or
=self-fertilization=. It will be seen that the =cross-fertilization=
relationship may be of many degrees--between two flowers in the same
cluster, between those in different clusters on the same branch,
between those on different plants. Usually fertilization takes place
only between plants of the same species or kind.
[Illustration: FIG. 195.--DIAGRAM TO REPRESENT FERTILIZATION.
_s_, stigma; _st_, style; _ov_, ovary; _o_, ovule; _p_, pollen-grain;
_pt_, pollen-tube; _e_, egg-cell; _m_, micropyle.]
In many cases there is, in effect, _an apparent selection of pollen_
when pollen from two or more sources is applied to the stigma.
Sometimes the foreign pollen, if from the same kind of plant, grows,
and fertilization results, while pollen from the same flower is less
promptly effective. If, however, no foreign pollen is present, the
pollen from the same flower may finally serve the same purpose.
In order that the pollen may grow, _the stigma must be ripe_. At this
stage the stigma is usually moist and sometimes sticky. A ripe stigma
is said to be =receptive=. The stigma may remain receptive for several
hours or even days, depending on the kind of plant, the weather, and
how soon pollen is received. Watch a certain flower every day to see
the anther locules open and the stigma ripen. When fertilization takes
place, the stigma dies. Observe, also, how soon the petals wither after
the stigma has received pollen.
=Pollination.=--The transfer of the pollen from anther to stigma is
known as =pollination=. The pollen may fall of its own weight on
the adjacent stigma, or it may be carried from flower to flower by
wind, insects, or other agents. There may be =self-pollination= or
=cross-pollination=, and of course it must always precede fertilization.
Usually the pollen is discharged by the bursting of the anthers. The
commonest method of discharge is through a _slit_ on either side of
the anther (Fig. 193). Sometimes it discharges through a _pore_ at the
apex, as in azalea (Fig. 196), rhododendron, huckleberry, wintergreen.
In some plants a part of the anther wall raises or falls as a _lid_,
as in barberry (Fig. 197), blue cohosh, May apple. The opening of an
anther (as also of a seed-pod) is known as =dehiscence= (_de_, from;
_hisco_, to gape). When an anther or seed pod opens, it is said to
_dehisce_.
[Illustration: FIG. 196.--ANTHER OF AZALEA, opening by terminal pores.]
[Illustration: FIG. 197.--BARBERRY STAMEN, with anther opening by lids.]
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