_Most flowers are so constructed as to increase the chances of
cross-pollination._ We have seen that the stigma may have the power
of choosing foreign pollen. The commonest means of necessitating
cross-pollination is the _different times of maturing of stamens and
pistils in the same flower_. In most cases the stamens mature first:
the flower is then =proterandrous=. When the pistils mature first, the
flower is =proterogynous=. (_Aner_, _andr_, is a Greek root often used,
in combinations, for stamen, and _gyne_ for pistil.) The difference
in time of ripening may be an hour or two, or it may be a day. The
ripening of the stamens and pistils at different times is known as
=dichogamy=, and flowers of such character are said to be dichogamous.
There is little chance for dichogamous flowers to pollinate themselves.
Many flowers are _imperfectly dichogamous_--some of the anthers
mature simultaneously with the pistils, so that there is chance for
self-pollination in case foreign pollen does not arrive. Even when
the stigma receives pollen from its own flower, cross-fertilization
may result. The hollyhock is proterandrous. Fig. 198 shows a flower
recently expanded. The center is occupied by the column of stamens. In
Fig. 199, showing an older flower, the long styles are conspicuous.
[Illustration: FIG. 198.--FLOWER OF HOLLYHOCK; proterandrous.]
[Illustration: FIG. 199.--OLDER FLOWER OF HOLLYHOCK.]
_Some flowers are so constructed as to prohibit self-pollination._
Very irregular flowers are usually of this kind. With some of them,
the petals form a sac to inclose the anthers and the pollen cannot be
shed on the stigma but is retained until a bee forces the sac open;
the pollen is rubbed on the hairs of the bee and transported. Regular
flowers usually depend mostly on dichogamy and the selective power
of the pistil to insure crossing. _Flowers that are very irregular
and provided with nectar and strong perfume are usually pollinated by
insects._ Gaudy colors probably attract insects in many cases, but
perfume appears to be a greater attraction.
[Illustration: FIG. 200.--FLOWER OF LARKSPUR.]
[Illustration: FIG. 201.--ENVELOPES OF A LARKSPUR. There are five wide
sepals, the upper one being spurred. There are four small petals.]
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