The rest of the chapter is devoted to a discussion of abortive and vestigial organs, whose existence Darwin naturally turns to great advantage in his argument for evolution. Throughout the whole chapter Darwin's preoccupation with the problems of classification is clearly manifest. On the question as to whether descent was monophyletic or polyphyletic Darwin expressed no dogmatic opinion. "I believe that animals have descended from at most only four or five progenitors, and plants from an equal or lesser number.... I should infer from analogy that probably all the organic beings which have ever lived on this earth have descended from one primordial form, into which life was first breathed" (p. 484). Darwin rightly laid much stress upon the morphological evidence for evolution,[356] which he considered to be weighty. It probably contributed greatly to the success of his theory. Though he himself did little or no work in pure morphology, he was alive to the importance of such work,[357] and followed with interest the progress of evolutionary morphology, incorporating some of its results in later editions of the _Origin_, and in his _Descent of Man_ (1871). In his morphology Darwin was hardly up to date. He does not seem to have known at first hand the splendid work of the German morphologists, such as Rathke and Reichert; he pays no attention to the cell-theory, nor to the germ-layer theory. His sources are, in the main, Geoffroy St Hilaire, Owen, von Baer, Agassiz, Milne-Edwards, and Huxley. Perhaps his greatest omission was that he did not give any adequate treatment of the problem of functional adaptation and the correlation of parts. It is not too much to say that Darwin not only disregarded these problems almost entirely, but by his insistence upon ecological adaptation and upon certain superficial aspects of correlation, succeeded in giving to the words "adaptation" and "correlation" a new signification, whereby they lost to a large extent their true and original functional meaning.