Form and Function: A Contribution to the History of Animal MorphologyRussell, E. S. (Edward Stuart)
History
Form and Function: A Contribution to the History of Animal Morphology
Russell, E. S. (Edward Stuart)
Anatomy, Comparative -- History; Morphology (Animals) -- History
It is true that Darwin himself, as well as his successors, believed that
natural selection was all-powerful to account for the evolution of the
most complicated organs, but it may be questioned whether he realised
all the conditions of the problem of which he thus easily disposed. He
says, rightly, in an important passage, that "It is generally
acknowledged that all organic beings have been formed on two great
laws--Unity of Type, and the Conditions of Existence. By unity of type
is meant that fundamental agreement in structure which we see in organic
beings of the same class, and which is quite independent of their habits
of life. On my theory, unity of type is explained by unity of descent.
The expression of conditions of existence, so often insisted upon by the
illustrious Cuvier, is fully embraced by the principle of natural
selection. For natural selection acts by either now adapting _the
varying parts of each being to its organic and inorganic conditions of
life_:[358] or by having adapted them during past periods of time: the
adaptations being aided in many cases by the increased use or disuse of
parts, being affected by the direct action of the external conditions of
life, and subjected in all cases to the several laws of growth and
variation. Hence, in fact, the law of the Conditions of Existence is the
higher law; as it includes, through the inheritance of former variations
and adaptations, that of Unity of Type" (_Origin_, 6th ed., Pop.
Impression, pp. 260-1). It is clear that Darwin took the phrase
"Conditions of Existence" to mean the environmental conditions, and the
law of the Conditions of Existence to mean the law of adaptation to
environment. But that is not what Cuvier meant by the phrase: he
understood by it the principle of the co-ordination of the parts to form
the whole, the essential condition for the existence of any organism
whatsoever (see above, Chap. III., p. 34).
Of this thought there is in Darwin little trace, and that is why he did
not sufficiently appreciate the weight of the argument brought against
his theory that it did not account for the correlation of variations.
Public-domain text, read in full here on John Shaqi.
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